Xelia Ermine Trace

prefix Ermine Trace · British Shorthair · line: Nadeya · the source records no eye colour · Variant known, this cat untested · also in its ancestry: Igor

Genealogy trace, one printable page carrying the proposed code

How this cat is connected to the trait

1. Xelia Ermine Trace, BRI ny 11
2. Nefertiti Ermine Trace, BSH ny 25 69, the dam of Xelia Ermine Trace
3. Ermine Trace Lemniscata, BLH ny 09 11 64, the dam of Nefertiti Ermine Trace
4. Ermine Trace Xrustal, BSH ny 11 33, the sire of Ermine Trace Lemniscata
5. Ermine Trace Nadeya, BSH ns 09 11 68, the dam of Ermine Trace Xrustal. genotyped DBE-NAD; a founder of this survey

That is 4 recorded parent links, and the animal at the end of it has a laboratory result, so what this cat could carry is not a guess about a line but a consequence of a test on a relative. Every step is a parent recorded in this register from a document; none of them is an assumption. The full genealogy is drawn below, and the sources for each animal are on its own page.

Save this cat as a PDF the connection, the genealogy and the source on one A4 landscape sheet, carrying this directory's name and address

The variant available to this cat

DBE-nad, not established for this animal. A DNA genotype sits 4 generations above this cat. Ermine Trace Nadeya is genotyped DBE-NAD. That does not settle what this cat carries, because a heterozygote passes the variant to half its offspring and the generations between are not tested. It names the variant available to this animal and the one to test for.

More than one variant could be present in this cat

DBE-NAD and Igor: PAX3, allele not published. More than one variant could be present in this cat: DBE-nad, Igor: PAX3, allele not published. It is not established that any of them is, and it is not established that two are. A dominant allele comes from a parent, and this cat's recorded ancestry carries an attribution of each. The attributions above it are on Ermine Trace Cherry, Ermine Trace Enya, Ermine Trace Jantar, Ermine Trace Jeana, Ermine Trace Kashemir, Ermine Trace Lemniscata, Ermine Trace Nadeya, Ermine Trace Olwen, Ermine Trace Qorall, Ermine Trace Sapphire, Ermine Trace Witch, Ermine Trace Xrustal, Jaki Ami Sissy Crazy Blue, Leonardo De Cassis, Nefertiti Ermine Trace, Schoenweg Teodoro. A cat holding two is a compound heterozygote, which this survey records as the state associated with deaf and dead kittens, and nothing but a test distinguishes one from two.

Computed from the recorded parentage at every build, counting both the variants this survey adopts and the ones it records a source claiming and does not adopt. It is an upper bound and not a finding: it says what could be here, never what is.

What the pedigree implies

Possible carrier by descent. DBE-NAD, through Ermine Trace Enya, Ermine Trace Jantar, Ermine Trace Lemniscata; the Igor line, PAX3, allele not published, through Jaki Ami Sissy Crazy Blue, Leonardo De Cassis; the Nadeya line, DBE-NAD, though no traced descent from the genotyped founder reaches this cat, through CH Carajan zur Sternebucht, CH Dellary's Exoti Merlin, CH Du Blue D'or Argentin Nuance's Golden

This is computed from the recorded parentage each time this site is built and is not typed into the register. It is a deduction from the trait being dominant, not a statement by any source, and it is not the same thing as the record's own status field. Only a laboratory settles it for an individual. The full list is on latent lines.

What is flagged on this tree

1. Ermine trace Enya. 68 AND 69 alternating down a single-source descent, in a second breed. On these five sheets the line runs Ermine trace Nadeya, BSH ns 09 11 68, to Ermine trace Enya, BSH ns 11 69, to Ermine trace Sapphire, BSH ns 09 12 68, to Ermine trace Jantar, BSH ny 09 25 68, to Nefertiti Ermine trace, BSH ny 25 69. Five generations and the code goes 68, 69, 68, 68, 69. The second branch does the same: Enya to Zarafina Ermine trace, BSH ny 11 68, to Faberge Ermine trace, BSH ny 09 11 68. Every sire brought in against these dams carries no eye code at all: Ermine Trace Kashemir BSH ns 11, Lusy-Mur Dionis BSH ay 11, Ermine Trace Qorall BSH ny 12, and no animal above any of them carries one either. There is no second source on these sheets. This is the Arst Venezia result reached again in a different breed and a different country: A single variant segregating in one line is sufficient to produce both codes. Nothing in this material has been genotyped
2. Ermine trace Qorall. The highest coefficients of inbreeding in this register, and the doubled cat carries no eye code. Smile Ermine Trace returns 25.0 PER cent on twenty-two unique ancestors of thirty, and Tanatos Ermine Trace returns 20.3 PER cent on nineteen of thirty. In both cases Ermine trace Qorall, BSH ny 12, stands twice: he is the sire of Smile and also the sire of Smile's dam Zarafina, which is a father to daughter mating; and he is the sire of Tanatos's sire Xrustal AND the sire of Tanatos's dam Zarafina, which makes those two half-siblings. Ermine trace Nadeya, the founder, also stands twice on the Tanatos sheet. The register records this as a fact about the matings and not as a judgement. What it shows is that the coded cats of this line are being produced by doubling back onto the line, which is what a breeder does to fix a trait, and that the animal doubled is one that carries no eye code at all
3. Ermine trace Nadeya. Attribution. The chart names this line's variant and this register had it as unidentified. Genealogy C13 is headed 'Nanotigr blue-eyed British shorthairs, DBE-ALT' and writes DBE-ALT in red beside Oliver, beside Amelia Nanotigr and beside Ermine trace Nadeya, who is the foundation cat of the Ermine Trace dominant blue-eyed line. The Ermine trace and Nanotigr cats are therefore attributed DBE-ALT by their own chart. It is an attribution and not a test: no Ermine Trace cat holds a laboratory result, and the only result ever held for Nadeya was a colourpoint test at the C locus, which this register corrected on 17 August 2026. The variant is recorded as claimed and not adopted, which is what the variant_claimed field is for, and the line stops being counted as a variant of its own | the variant is named and she is the cat it is named for, 12 September 2026. Abitbol M, Eisele T and Lipovenko E, 'Additional PAX3 Variants for Dominant Blue Eyes in Cats', Animal Genetics 2026;57:e70199, doi:10.1002/age.70199, open access, read 12 September 2026 names the allele DBE-NAD, Nadeya Dominant Blue Eyes, NC_058375.1:g.[205097674_207411974del;207411975_211290497inv;211290498_211387174del]. It is not a point change but a complex rearrangement, a deletion then an inversion then a second deletion, spanning about 6.3 million bases and taking PAX3 with it. She is recorded heterozygous. All 39 DBE cats of the line were heterozygous, all 47 controls of the line and breed were wild-type, and the allele was absent from every DBE cat of every other line tested. No Living homozygote has ever been found, which is the basis for the paper's statement that the homozygous genotype is likely lethal. The DBE-ALT attribution from chart C13 is withdrawn by this result and is no longer carried in any field: the chart said DBE-ALT, the genotype says DBE-NAD, and the genotype settles it. The chart remains named here as the source that said otherwise. Her parents were two registered British Shorthairs and neither was a dominant blue eye: the paper attributes the sire's blue eyes to colourpoint and gives the dam as green-eyed. S_PAX3_2026_234 | an english-language note on this line is in circulation and it is wrong. A 2026 update added to the translation of the cattery's own 2021 article states that testing has shown the Ermine Trace cats to carry DBE-ALT, and goes on to infer that DBE-ALT may be deeply embedded in the silver and golden British population and that cats registered 61 are more likely DBE-ALT. The published genotyping says the opposite. The line carries DBE-NAD, an allele found in no other line tested; all 39 dominant blue eyed cats of the line are heterozygous for it; and DBE-ALT is excluded, two Ermine Trace cats having already been genotyped wild-type for the DBE-CEL insertion in 2024 and the line being wild-type for DBE-ALT throughout. The inference drawn from the error is therefore unsupported too: nothing in the published work places DBE-ALT in the wider silver and golden British population on the strength of this line. The note is recorded here because a breeder reading it in English would otherwise act on it. Source: the Ermine Trace cattery's own published account of the line, 'Reviewing the dominant blue (DBE) gene in the Ermine Trace cattery', 2021, in English translation. A breeder statement and not a peer-reviewed source, and the counts in it are the cattery's own. S_ERMINE_238 | the one place this line meets another DBE line, and it is not a route for either variant. Oliver, the founder of the Nanotigr line, and this cat share 55 recorded ancestors, among them Eric von Buergersruh, who stands above both: he is the sire of Captain Ermine Trace, Oliver's great-great-grandsire, and he is Ermine Trace Olwen's great-grandsire. The two lines carry different published alleles: Nanotigr is DBE-ALT and this line is DBE-NAD, and neither allele appears in the other line on any test. The shared ancestry therefore cannot be the route for either, and 55 ancestors in common says only that both are bred from the same British chinchilla stock, which every British chinchilla cattery in Europe is. It is the clearest worked example this register holds of its own rule: shared ancestry is not a shared variant, and here two genotypes prove it rather than merely warn about it. Eric von Buergersruh was a widely-used sire, which is what makes him a common ancestor and not a source. S_ERMINE_238 | the Igor line does not stand above this cat, and it does stand below her. The founding tree of this line, 179 animals counting everything above Nadeya and above Oliver of the Nanotigr line together, contains not one record carried under an Igor label. Nothing of the Igor material is ancestral to her, which removes the only route by which her determinant could have arrived from that line and leaves the cattery's own account, a variant arising in her, standing. Below her it is the other way round: of the 30 cats that descend from her by unbroken parent links, 28 also have an ancestor carried under the Igor label, every one of those paths running through Leonardo De Cassis and Jaki Ami Sissy Crazy Blue, both of whom this register holds with no eye colour and no variant. The two halves say different things. Above her, the absence is evidence about where DBE-NAD came from. Below her, the presence is a compound heterozygote question and not an origin question, and one genotype on either of those two animals settles it. S_ERMINE_238 | the cattery has restated its own codes and the register holds the older form. Its public site, read 13 September 2026, gives this cat as BRI ns 11 08 61 where this register holds BSH ns 09 11 68, and the same substitution runs through its other cats: Barkhat as BRI as 11 33 08 against BSH as 09 11 33 68 here, Faberge as BRI ny 11 08 61 against BSH ny 09 11 68. The cattery is writing 08 where this register has 09, AND 61 where it has 68. Both pairs are eye and white markers in the same family of codes and the substitution is systematic rather than a slip. The register does not change its codes on a website reading: the codes held here come from pedigree sheets and certificates, and which form a registry recognises is the question case study one is about. Recorded so that the difference is not mistaken for an error in either source. S_ERMINE_238 | asked whether the determinant sits above her, the register answers from its own parent links. If cats carrying DBE-NAD existed that did not descend from her, it would have come from one of her parents. Not one does. Every record in this register that carries DBE-NAD descends from her by unbroken parent links. Her sire ch Schoenweg Teodoro has one other recorded offspring here, Ermine trace Kashemir, BSH ns 11, and Kashemir carries no eye code and no 33 to hide one, so his state is readable and he shows nothing. Her dam Ermine Trace Olwen, BSH ns 11, is likewise readable and shows nothing, and the cattery records that Olwen's own dam produced five kittens in three litters with nothing unusual in any of the others. The one animal above her whose state cannot be read is Teodoro himself, a colourpoint at ns 11 33, and the cattery counts about forty offspring by him before her, gold and silver, colourpoint and chinchilla, every one normal. A dominant determinant carried by a sire shows in about half his kittens and one in forty is not half. The determinant appears in her and not above her on everything this register holds. No cat showing the trait descends from Teodoro or from Olwen without also descending from her. S_ABOVE_247 | the question of whether the determinant sits above her is closed by a count of her sire's litters. CH Schoenweg Teodoro sired thirteen litters and she is the only dominant blue eye kitten in any of them. A dominant reaches about half of every litter, so a carrier sire would have shown it in nearly all thirteen. He did not carry it. Her dam is readable and shows nothing, her dam's dam produced five kittens in three litters with nothing unusual in any other, and every record in this register carrying DBE-NAD descends from her. The mutation arose in her. S_TEODORO_248 | she is a founder in the strict sense and not merely the earliest cat held. Both her parents' pedigrees are now held in full and both are empty of the trait: her sire gave thirteen litters with one dominant blue eye kitten in all of them, and her dam returns thirty ancestors over four complete generations with no eye code on any of them. The variant arose in her. That is what the published work reports as well, DBE-NAD being found in no other line tested. S_OLWEN_249 | the test that would break the founder reading was run on 13 SEPTEMBER 2026 and it did not break it. If the determinant came from a parent rather than arising in her, cats carrying it would exist that were born before her or that descend from something other than her. The register holds thirty-eight records carrying DBE-nad besides herself. Not one was born on or before 12 OCTOBER 2016. Not one fails to descend from her by unbroken parent links. The earliest is 10 November 2017, thirteen months after her. Two of the thirty-eight carry no date at all, Aquarelle Ermine Trace and Elina Ermine Trace, and both descend from her, so neither is a counter-example either. Twenty-four Ermine trace litters are now held with a date and every kitten in every one of them reaches her. S_LITTERS_250 | the timeline was tested on 13 SEPTEMBER 2026 and it holds at every step. A descent is only real if each cat is old enough to have bred when the next was born, and if each generation sits far enough below the one above it in time. Ninety-two records descend from her and one hundred and forty-three parent to kitten pairs carry a date on both sides. In every one of the 143 the parent is older by at least 300 DAYS. Not one kitten is born on or before a parent, and not one parent is too Young to have bred. The generations also sit where they should. Her own first generation begins 13 months after her, the second at 32 months, the third at 48, the fourth at 73 and the fifth at 101. The shortest possible spacing for five generations is about 50 months and the record uses 101, so the descent is not compressed against the calendar anywhere. Four records in the descent carry no date. S_LITTERS_250 | with the Emerald Flower branch re-lined on 13 SEPTEMBER 2026, no cat anywhere in this register shows the trait under the Nadeya label without descending from her. Wanessa and Paula Emerald Flower were the only two, they were labelled from a breeder chart, and Wanessa's own pedigree carries no Ermine Trace cat and no eye code on any of its twenty-eight ancestors. S_WANESSA_252 | the route by which the Emerald Flower branch touched this line has been traced and it carries nothing. Two animals appear in both bodies of material, Vitma Belle Gold and IC Silver Angel's Jantar, at the great-great-grandparent rank. Both pedigrees are now held and neither carries an eye code on any of its forty combined ancestors. S_SHARED_253 | the ALT material and this line do not meet by descent. No record in this register descends from both her and a DBE-ALT cat, no DBE-ALT cat stands below her, and none stands above her. The two variants share trace ancestors in the golden British foundation stock and nothing else. S_SHARED_253 | where this variant sits in PAX3. A third party compilation places DBE-NAD at not a point variant at all but a large structural one of about 6.29 megabases spanning the PAX3 REGION, described as a deletion, an inversion and a deletion. The register records the position and does not adopt it as its own finding: a schematic on a website is not a paper, and where this register needs the science it cites the papers directly. It is recorded because it is checkable, and because the position in the gene is the one thing about these variants that a breeder can never observe and a laboratory never reports on a result sheet. S_LOCI_272 | an outside count that agrees with this one. The same third party compilation holds eight named variants, DBE-CEL, DBE-ALT, DBE-re, DBE-AGO, DBE-Maru, DBE-nad, DBE-cDc and DBE-xHW, and states that six of them are characterised in peer-reviewed research. That is this register's position exactly: eight held, six published, and the two Chinese ones identified in 2025 but not published. Two compilations working from the same Literature and arriving at the same eight, and the same split between them, is worth recording because the count is the figure this survey is most often asked for. S_LOCI_272 | the split between the eight is not six identified and two unknown. It is six characterised in peer-reviewed research and two identified by a commercial laboratory that does not publish, with a test on sale for each of those two. The distinction matters to a breeder, who can buy a test for all eight of the identified variants and for none of the unidentified ones. S_CHINESE_273
4. Ermine trace Sapphire. 68 AND 69 alternating down a single-source descent, in a second breed. On these five sheets the line runs Ermine trace Nadeya, BSH ns 09 11 68, to Ermine trace Enya, BSH ns 11 69, to Ermine trace Sapphire, BSH ns 09 12 68, to Ermine trace Jantar, BSH ny 09 25 68, to Nefertiti Ermine trace, BSH ny 25 69. Five generations and the code goes 68, 69, 68, 68, 69. The second branch does the same: Enya to Zarafina Ermine trace, BSH ny 11 68, to Faberge Ermine trace, BSH ny 09 11 68. Every sire brought in against these dams carries no eye code at all: Ermine Trace Kashemir BSH ns 11, Lusy-Mur Dionis BSH ay 11, Ermine Trace Qorall BSH ny 12, and no animal above any of them carries one either. There is no second source on these sheets. This is the Arst Venezia result reached again in a different breed and a different country: A single variant segregating in one line is sufficient to produce both codes. Nothing in this material has been genotyped
5. Ermine trace Xrustal. The code goes 68, 69, 68 and crosses into a second breed. Unreal ZouWu Ermine Trace is BLH ny 11 68, a British longhair, out of Urbi et Orbi Ermine trace, BSH ns 09 11 69, out of Ermine trace Xrustal and Ermine trace Nadeya, BSH ns 09 11 68. His dam Manila Ermine trace is BLH ny 12 with no eye code, and neither she nor any of the fourteen Boon animals above her carries one. THE 68 on him came from a 69 sire and from nowhere else | he carries no white at all, ny 11 being black golden shaded, where his sire carries 09. The white drops out of the code in one generation and the eye does not | Ermine trace Xrustal now has four coded descendants and he was placed as not carrying the gene: Lemniscata 64, Tanatos 68, Urbi et Orbi 69, and through Urbi et Orbi this grandson at 68 in another breed | no variant is adopted for this cat and descent is not enough to adopt one. It descends from Ermine Trace Nadeya, who is genotyped DBE-NAD, but it shows no blue or odd eye and holds no test. A dominant determinant passes to about half the kittens of a carrier, so a descendant without the eye is as likely to be wild-type as to be a latent carrier. The register held this cat as DBE-nad between 12 AND 13 SEPTEMBER 2026 in error, on descent alone, and the attribution is withdrawn. One test on this animal settles it. S_ERMINE_238 | the variant is restored to him, and on a different basis from descent. He is an obligate carrier. His daughter Ermine Trace Lemniscata carries the determinant and is recorded latent; her other parent, Ermine Trace Witch, is not descended from Nadeya and does not carry it. A dominant allele cannot skip a generation, so it reached Lemniscata through this cat and through no one else. The cattery says the same in its own words, calling him a DBE heterozygote and recording that his daughter's latency took it by surprise because nothing in his appearance showed it. He is a colourpoint, ny 11 33, which is why nothing showed. S_ERMINE_238
6. Ermine trace Jantar. Confirmed. A chart reading now has a registry pedigree behind it. This register entered his sire and dam from the colour of a line on breeder-compiled chart C13 and recorded that it had done so. The Pawpeds pedigree read 18 AUGUST 2026 gives the same sire and the same dam, so the parentage no longer rests on a chart
7. Ermine trace Lemniscata. The cat the breeder placed as clear, and what he has since produced. Ermine Trace Xrustal was placed with a cattery on the breeder's own account that he had not inherited the gene. He carries no eye code and he is BSH ny 11 33, a colourpoint, so a blue eye on him could not have been read as evidence of anything in any case. Registry sheets now give him two coded offspring: Ermine trace Lemniscata, BLH ny 09 11 64, whose daughter Nefertiti Ermine trace is BSH ny 25 69; and Tanatos Ermine trace, BSH ny 09 11 68. A cat believed clear sired a 68 and a 64 whose daughter is 69. This is what a latent carrier does, and it is the clearest demonstration in this register that an eye code is not a carrier test. Nothing here is genotyped, and the placement decision was made on appearance because no test was used | she keeps the variant where six of her relatives lost it, and the reason is the latency. Six descendants of Nadeya with no blue or odd eye were credited with DBE-NAD on descent alone and the attribution was withdrawn on 13 September 2026. This cat is different: the cattery records her as a LATENT, a carrier whose eye did not stay blue, and states that her eyes glowed red in kittenhood and then stopped. That is a positive statement that she carries it, not an inference from descent. It is still not a test. S_ERMINE_238
8. Nefertiti Ermine trace. 68 AND 69 alternating down a single-source descent, in a second breed. On these five sheets the line runs Ermine trace Nadeya, BSH ns 09 11 68, to Ermine trace Enya, BSH ns 11 69, to Ermine trace Sapphire, BSH ns 09 12 68, to Ermine trace Jantar, BSH ny 09 25 68, to Nefertiti Ermine trace, BSH ny 25 69. Five generations and the code goes 68, 69, 68, 68, 69. The second branch does the same: Enya to Zarafina Ermine trace, BSH ny 11 68, to Faberge Ermine trace, BSH ny 09 11 68. Every sire brought in against these dams carries no eye code at all: Ermine Trace Kashemir BSH ns 11, Lusy-Mur Dionis BSH ay 11, Ermine Trace Qorall BSH ny 12, and no animal above any of them carries one either. There is no second source on these sheets. This is the Arst Venezia result reached again in a different breed and a different country: A single variant segregating in one line is sufficient to produce both codes. Nothing in this material has been genotyped
9. Xelia Ermine trace. The sire is named and not entered. The sheet gives him as Virtuoso Wizard MaxelPoint, BRI ay 12, and this register holds no record for him, so the sire side of this litter is empty. S_ERMINE_LITTERS_239

A flag is not an error. It marks a place where two sources say different things, or where an attribution cannot be made on the evidence held.

The record

Register id

erm-0110

Sex

not stated by the source

Cattery prefix

Ermine Trace

Breed, as stated

British Shorthair

Line

Nadeya

Eye status

unknown: the source records no eye colour

Role

offspring

Litter key

Ermine Trace X6, 8 December 2023

Year

8 December 2023

EMS code

BRI ny 11

The code read out

British; black golden; shaded

The description held

No eye code on the cattery sheet, so nothing is established about its eyes. No variant is adopted.

Note

The code and the description do not use the same words for this coat. Both are recorded and neither is reconciled to the other.

Evidence grade

Reported asserted, with neither a test nor a traced descent

In the register

• Parents: Nefertiti Ermine Trace
• Littermates (Ermine Trace X6, 8 December 2023): Xaos Ermine Trace, Xeon Ermine Trace, Xoxloma Ermine Trace

This directory is a working aid compiled from published charts, registry documents and breeder statements. It is not a register and it has no standing of any kind. The grade attaches to the animal, not to the line: a cat whose ancestor was genotyped is line-traced, not genotyped.