Variant trace

What each variant reaches through recorded parent links alone, with cattery prefixes ignored, recomputed on every build

The material on this page is drawn from the Global Cat Health Survey 2026, which is where its sources and its declarations of interest are set out in full.

Every line this register holds, and how much ancestry each pair shares. Circle size is the number of recorded ancestors. The number on a line is the ancestors the two have in common. Sharing ancestors is not carrying a variant: where a variant has actually reached another line, it is set out under shared lineal descent below.

Shared ancestors between every pair of lines Three different PAX3 variants, not one 470 343 159 62 156 139 158 158 3 DBE-ALT 1,117 Igor 767 DBE-CEL 392 DBE-NAD 361 DBE-RE 327 DBE-MARU 17, no links Circle size is the number of recorded ancestors. Line weight and the number on it are the ancestors the two share. The ringed three are three different PAX3 variants. Genotyping separates them. The case is set out in the Igor case study.

What this page reports is shared ancestry, not direction. Where a variant is shown to share ancestors with the Igor chart, that means the two pedigrees meet at a common ancestor. It does not mean the variant entered the Igor lines, and it does not mean the Igor line entered the variant's. Both directions have been tested and the pedigrees held exclude both: neither population is ancestral to the other. New pedigrees could change the picture in either direction.

Each variant, when its founder appeared and when it was named

A variant is named years after the cat it is named for was bred from. In every case here the founder was producing kittens, and those kittens were being sold and bred on, long before any laboratory could say what they carried. The bar is that interval, and it is the only measure the survey can give of how long a population goes untestable.

Each variant, when its founder appeared and when it was named Every one of these cats was bred from for years before anyone could name what it carried The bar is the interval in which a breeder had no test. Founder dates are this register's; naming dates are the publications. 19952000 20102015 20202026 DBE-ALT Fyodor, Kazakhstan 29 years DBE-CEL Roxi, Kazakhstan about 26 years DBE-RE Rociri Elvis, Netherlands 9 years DBE-NAD Ermine Trace Nadeya, Russia 10 years DBE-MARU Marusja, Russia 7 years DBE-AGO Agostino, Maine Coon named 2025, founder date not recorded here DBE-cDc, DBE-xHW China identified 2025, unpublished The Igor determinant Igor, Kazakhstan still running Twenty-nine years for the first, seven for the most recent. The interval is shortening and it has not closed.

The interval is shortening and it has not closed. Twenty-nine years separated the first founder from its allele and seven separated the most recent. The Igor line's bar is still running, and it is the largest population in this register without a published allele. Nothing about the trend makes the gap safe: a cat bred during the interval is bred blind, and every cat bred out of it inherits that.

Every variant this register holds, named and unidentified

A blue eye that is already explained is not on this page. Where a cat's own code carries a pointing factor, 31 Burmese, 32 mink or 33 colourpoint, the pointed gene is an ordinary cause of its blue eye, and where a whole breed group is of the colourpoint series it is not a group of unknown variant unless a known variant has been bred into it. One chart in this register, thirty-six cats headed Dominant Blue Eyes in Mink Ragdoll, is excluded on that ground and is counted nowhere below. The rule is set out on the method page.

The eight identified variants are at PAX3 and all eight are dominant, so one copy is enough to show the eye and a carrier that does not show it cannot be identified without a test. The unidentified ones are taken to be dominant on the same basis, because they produce the same phenotype in the same pattern; that is an assumption. Attributed counts every register entry to which some source gives that variant, at any evidence grade; for an unidentified variant it counts the cats of the line that show a blue or odd eye. DNA counts those with a laboratory result.

VariantStatusNamed forTraced toOriginAttributedDNA
DBE-CELidentifiedCeleste, also written CelestialRoxi, also written Roksi and Roxy DarlinKazakhstan241
DBE-ALTidentifiedthe AltaiFyodor and Nikita'l Seymour, two cats carrying itUst-Kamenogorsk, Kazakhstan, 1995 or 19964816
DBE-REidentifiedRociri Elvis, a Maine Coon born in the Netherlands on 7 May 2015Rociri Elvis himselfNetherlands3012
DBE-AGOidentifiedAgostino, a Maine CoonAgostino himselfnot recorded here1none
DBE-cDcidentifiednot recorded hereno cat at allChina2none
DBE-xHWidentifiedXin-He Winnie, a Chinese domestic catXin-He Winnie herselfChina4none
DBE-NADidentifiedErmine Trace Nadeya, a British Shorthair born in Russia on 12 October 2016Nadeya herself, heterozygousRussia421
DBE-MARUidentifiedMarusja DBE, an outbred domestic shorthair, line founded in Russia in 2019Marusya herself, heterozygousRussia, Volgograd51
Igor: PAX3, allele not publishedunidentifiedno allele namedthe line itself, 400 records60 cattery prefixes71none
Blue Eyed Maine Coons (Markiss), DBE-CEL by test on the generation belowidentified, no cat of it testedDBE-CEL, by published genotyping on the generation below the founding mating, which also excluded DBE-ALTthe line itself, 98 records19 cattery prefixes54none
Siberian, Altai or Barnaul unconfirmed, unidentifiedunidentifiedno allele namedthe line itself, 130 records35 cattery prefixes51none
Topaz foundation (C6 and C11), unidentifiedunidentifiedno allele namedthe line itself, 57 records7 cattery prefixes20none
Karagaisky, unidentifiedunidentifiedno allele namedthe line itself, 18 records2 cattery prefixes14none
Highlander, Olynx (C11), unidentifiedunidentifiedno allele namedthe line itself, 10 records2 cattery prefixes5none
King's Way, unidentifiedunidentifiedno allele namedthe line itself, 7 records2 cattery prefixes5none
Pillowtalk-Foreigner, unidentifiedunidentifiedno allele namedthe line itself, 61 records18 cattery prefixes3none
Royal Edelweiss, unidentifiedunidentifiedno allele namedthe line itself, 2 records1 cattery prefix1none
Gosha, unidentifiedunidentifiedno allele namedthe line itself, 1 record0 cattery prefixes0none

The eight identified variants have been sequenced and named. The 10 unidentified ones below them are documented breeding programmes in which a dominant blue eye segregates and no animal has been genotyped. They are traced here on the same basis as the named six, since a cat carrying an unnamed allele can produce a compound heterozygote as readily as a cat carrying a named one.

DBE-CEL. The first of the eight to be identified and published. The LOOF requirement of 2019 is that every Celeste descend from Lazuli, who is the one animal in this register with a laboratory result for it.

DBE-ALT. Published alongside DBE-RE, which separated the two Kazakh variants. It is the variant with the most laboratory results in this register.

DBE-RE. PAX3 c.937C>T, a nonsense variant. The cat it is named for is coded 68 on a certified pedigree, and no laboratory result for that cat himself has been published or supplied, so his own grade in this register is reported.

DBE-AGO. PAX3 c.160del, a frameshift in exon 2, published June 2025. The combination of DBE-RE and DBE-AGO is the proposed compound heterozygote risk case in the Maine Coon.

DBE-cDc. Named August 2025 and since detected in the Abyssinian within that population. It is the one variant of the eight with no named cat anywhere in this register.

DBE-xHW. Named September 2025 and unpublished. It is the only one of the eight with no code on any commercial panel: the laboratory that identified it sells a panel of five and this is not among them, so it is identified by exclusion and not by test.

DBE-NAD. Published 2026 and the largest of the eight. Not a point change but a complex rearrangement: a deletion, then an inversion, then a second deletion, spanning about 6.3 million bases and taking PAX3 with it. NC_058375.1:g.[205097674_207411974del;207411975_211290497inv;211290498_211387174del]. All 39 DBE cats of the line were heterozygous for it and all 47 controls were wild-type. No cat of any other line carries it.

DBE-MARU. Published 2026. PAX3 XM_019838731.3:c.784C>T, a nonsense variant in the fifth exon, predicted to truncate the protein at p.(Arg262*) and so to remove 46 per cent of it. All 14 DBE cats of the line were heterozygous. The one animal recorded homozygous was an abnormal kitten, dead at birth, out of two DBE parents.

Igor: PAX3, allele not published. PAX3, allele not published. Three published rounds of genotyping place the determinant at PAX3 by excluding four named alleles from this line. What is absent is a published genotype, not an identification. Four register labels, one breeding programme. Eight PawPeds sheets read 18 August 2026 put the founder in the same place with all four positions above him empty, and a second parentless coded founder, BG'Amur Dream, stands behind the Kittystar cats. Its compiler states that it reached seven named British Shorthair and Longhair catteries, SuperQueen, King Pride, Kittystar, SoloMiya, Lana's Story, Helprada and Beauty Factory, and that the founder probably carries the Altai gene, which is unconfirmed. No Altai cat is named in any account, so no pedigree can settle it. Two further statements are held and both are queried: that DBE-ALT is present in some Igor lines and it is not known which, and that cats of one cattery in this line were tested and found clear of DBE-ALT, DBE-RE and DBE-CEL. Neither names an animal, so nothing is built on either.

Blue Eyed Maine Coons (Markiss), DBE-CEL by test on the generation below. The allele this line carries is not unidentified. It is DBE-CEL, and a laboratory says so. The line begins in the Roxi by Seymour outcross into the Maine Coon, made twice over two unrelated queens, so the founding mating held one parent of each variant and either could have passed. The published genotyping on the generation below it settles which did: ten Topaz Maine Coons returned wild type for DBE-ALT and nine returned heterozygous for DBE-CEL. DBE-ALT is therefore excluded by test and DBE-CEL is carried. What has not been tested is the animals themselves. Both named offspring carry eye code 68 and neither has a result of its own, so every cat of this line is line-traced to DBE-CEL rather than genotyped for it, and it is listed here for that reason and not because the variant is unknown.

Siberian, Altai or Barnaul unconfirmed, unidentified. The largest untested population in the register by cattery count.

Topaz foundation (C6 and C11), unidentified. The founder population both Kazakh variants come out of. Neither founder has been genotyped; the variants are attributed to their lines through tested descendants.

Karagaisky, unidentified. The Munchkin line, from a village domestic cat at Chelyabinsk. Every coded cat in it also carries white spotting.

Pillowtalk-Foreigner, unidentified. The only line here that has been tested. A cat of it showed the eye and was clear of DBE-ALT, DBE-RE and DBE-CEL, which were the three variants described at the time. Retesting one cat against a current panel would either confirm a ninth variant or resolve the line to DBE-AGO. It is a German line, named for Venezia the Foreigner, and is reported restricted to the Maine Coon to date, which the register agrees with: all 61 records are Maine Coon. There is no data on homozygosity and none on deafness, and the search for the variant is reported ongoing by a German research team. Abitbol M, presentation slides, 2026, supplied to the survey. Reported: the statement carries no genotype.

Gosha, unidentified. A Russian line, its founder registered XSH with no further information recorded, and reported introduced into the Devon Rex and the Munchkin. There is no data on homozygosity and none on deafness, and the search for the variant is reported to have just started. The register holds one cat of this line and no descent from it, so neither introduction can be followed animal by animal here; the Devon Rex is carried in the breed table on the Laboratories page and no Devon Rex is entered in the register. Abitbol M, presentation slides, 2026, supplied to the survey; founder from messybeast.com. Reported: neither statement carries a genotype.

The cats with a determinant from each parent

A compound heterozygote is one copy each of two different variants, and it is the state in which harm is documented in this trait. A cat cannot be one unless a determinant came down each side. This is that list, and it is the shortest and the most important on this page: 15 cats, computed from parent links at every build.

CatBornSire carriesDam carriesTested
Solomiya Josephine1 October 2022IGORALTno
Amery Dolce Blu12 April 2024REALT, CELno
Miyuki of Aerlin, given on the Celeste charts as Miyuki2016ALTCELno
Mochi2016ALTCELno
Nikita'l Markiss [Cherkassy, UA]26 March 2016ALTCELno
Dream Dolce Blu27 April 2023ALT, CELREno
Crishell Dolce Blu31 January 2024REALT, CELno
Abigail Dolce Blu8 April 2024REALT, CELno
Mavi of Aerlinborn 18 July 2016ALTCELno
Mistralnot heldALTCELno
Merlinnot heldALTCELno
Matchanot heldALTCELno
Mavinot heldALTCELno
Maidi, printed on the chart as Maidi with a diaeresisnot heldALTCELno
the second daughter of the two-line cross, not named by the sourcenot heldIGORALTno

Not a list of compound heterozygotes. Each parent passes one allele of the two it holds, so a cat here may carry the variant from neither side, from one, or from both. Nothing but a genotype distinguishes the three. Every cat on this list is where a test would answer a question that cannot be answered any other way.

These are the first generation. Every cat below them inherits the same open question in a weaker form, and the whole population in which two variants could be present is listed further down this page under every cat in which two variants could be present. The 15 here are the matings that made that population, and they are where a test is worth most.

Shared lineal descent between the variants

A cat is in a variant's trace if it descends from that variant's founder by unbroken parent links, or if the register attributes that variant to the cat itself. Everything below either is in the trace as well, because a cat below a carrier could have inherited from it.

A cat mated into a trace is never added to it. Where a carrier is bred to an outside cat, that cat counts to its own trace and to nothing else. Only the kittens count to both, because only they can have inherited from either side. This is descent and not association: nothing on this table is computed upward, and shared ancestry, which is, is reported further down the page and measures common breed stock rather than the passage of an allele.

The second rule carries most of the weight. DBE-ALT is attributed to 45 animals of this register and 43 of them descend from Nikita'l Seymour by no recorded link at all. Sixteen of those 43 carry a laboratory genotype, and they are the whole of the DBE-ALT genotyped population: not one animal with a DBE-ALT result descends from the founder the variant is traced to. A trace drawn from the founder alone holds 104 cats. Drawn from the founder and the attributions together it holds 170. The founder is the earliest carrier this register can name, not the origin of the allele.

Attribution is not all one grade. A seed may be genotyped, line-traced or reported, and the grade is on each cat's own page. A trace is the population in which that allele could be present, computed from parentage, and it is not a list of cats that carry it.

TraceFounderStatusFrom the founderSeeded on an attributionIn the trace
DBE-ALTNikita'l Seymouridentified, PAX310443170
DBE-CELRoxiidentified, PAX31328140
NadeyaErmine Trace Nadeyaidentified, PAX392none92
IgorIgor Azure DreamPAX3, allele not published62none62
DBE-RERociri's Elvisidentified, PAX353154
DBE-MARUMarusja DBEidentified, PAX35none5
Schasti DomaMeowrigi Irisano variant identified4none4

DBE-AGO and DBE-xHW hold one reference animal each and no descent, so they cannot meet anything in this register. DBE-cDc is held by no animal at all. Their absence below is a gap in the record and not a finding of separation.

Where two traces meet

A variant has reached another line when it traces into it, meaning an animal stands in both traces. That is the first column and it is the measure. The second column is forward-looking and is not a trace: it counts the catteries holding cats of both, which is where a mating could still be made. Catteries are counted by prefix and a cat entered without one cannot be counted, so that column is a floor rather than a total.

PairAnimals in both tracesCatteries holding bothThe catteries
DBE-ALT and DBE-CEL8410arst, blueridge, daerlin, dolceblu, hanforsworld, hippopotamia, legendedesalpes, nikital, solareclipce, vomleineufer
DBE-ALT and DBE-RE53arst, belleayr, dolceblu
DBE-CEL and DBE-RE52arst, dolceblu
DBE-ALT and Igor23fromsham, solomiya, superqueen
DBE-ALT and Nadeya00none
DBE-ALT and DBE-MARU00none
DBE-ALT and Schasti Doma00none
DBE-CEL and Nadeya00none
DBE-CEL and Igor00none
DBE-CEL and DBE-MARU00none
DBE-CEL and Schasti Doma00none
DBE-MARU and Schasti Doma00none
DBE-RE and DBE-MARU00none
DBE-RE and Schasti Doma00none
Igor and DBE-RE00none
Igor and DBE-MARU00none
Igor and Schasti Doma00none
Nadeya and Igor00none
Nadeya and DBE-RE00none
Nadeya and DBE-MARU00none
Nadeya and Schasti Doma00none
The DBE-ALT, DBE-CEL and DBE-RE traces and their overlaps DBE-ALT 169 DBE-CEL, Roxi 140 DBE-RE 54 85 56 49 79 0 0 5 Igor 57 2 below both Areas are not to scale. The 5 in the centre are Maine Coons of one cattery, standing in all three lines.

The three named variants that have met each other. The 5 in the centre stand in all three traces at once, and they are five Maine Coon queens of one cattery. Areas are not to scale.

DBE-CEL by breed, split by whether the cat is in the registered Céleste line in the registered Céleste line, below Lazuli carries DBE-CEL outside it Maine Coon 55 Céleste 39 German Longhair 21 Siberian 7 Topaz 5 British Shorthair, British Longhair 4 Experimental Longhair 3 Lykoi 3 Domestic 2 Minuet 1 72 inside the registered line. 68 outside it, and 55 of those are Maine Coons. Lazuli, the cat every Céleste must descend from, is himself out of a Maine Coon.

DBE-CEL by breed, split by whether the cat is in the registered Céleste line. Lazuli is the animal the LOOF requires every Céleste to descend from, and he is himself out of a Maine Coon. The variant was in the Maine Coon before it was registered to the British side.

Read the first two columns together. A pair with animals in both traces has already met. A pair with none in both traces but catteries holding both has not met and could. A pair with neither cannot meet without a cat moving between catteries. The prohibition this directory proposes applies to the second class as much as the first, because that is the one where the mating has not happened yet.

The worked case, DBE-ALT against the Igor line, is set out in the Igor case study.

Where two variants meet, and which cats could carry both

The same question asked of each pair: did the second variant come from above the founder, in which case one descends from the other and it is an origin question, or only from below, in which case the two met in a later mating and the cats beneath them may hold either variant or both.

Where DBE-ALT and DBE-CEL meet ABOVE THE FOUNDER, WHERE THE VARIANT CAME FROM 142 animals above the DBE-ALT founders. 0 of them belong to DBE-CEL. DBE-ALT and DBE-CEL are not related by descent. This is the origin question, and it is closed. DBE-ALT in the genealogy: Pushok, Eliza, Nikita'l Seymour, Oliver genotyped: Cotomodda Hubba-Bubba BELOW THE FOUNDER, WHERE THE TWO LINES MEET 142 animals below the DBE-ALT founders. 84 also have DBE-CEL in the ancestry. DBE-CEL arrives through the other parent in a later mating, not down through these founders. All 84 come down from Pushok and Eliza. No cat in the Oliver branch is among them. the descent DBE-CEL enters by a later cross, and reaches these cats Merlin Nuvola and 82 more 84 cats stand below the collision of DBE-ALT and DBE-CEL A kitten below the collision can inherit DBE-ALT, or DBE-CEL, or BOTH VARIANTS IN ONE CAT, or NEITHER. Descent is not a diagnosis. Standing below two founding descents is not carrying anything, and only a test settles which of the four a given cat is. Carrying both is the compound state, and it is the one the survey records as associated with deaf and dead kittens.
Where DBE-ALT and DBE-RE meet ABOVE THE FOUNDER, WHERE THE VARIANT CAME FROM 142 animals above the DBE-ALT founders. 0 of them belong to DBE-RE. DBE-ALT and DBE-RE are not related by descent. This is the origin question, and it is closed. DBE-ALT in the genealogy: Pushok, Eliza, Nikita'l Seymour, Oliver genotyped: Cotomodda Hubba-Bubba BELOW THE FOUNDER, WHERE THE TWO LINES MEET 142 animals below the DBE-ALT founders. 5 also have DBE-RE in the ancestry. DBE-RE arrives through the other parent in a later mating, not down through these founders. All 5 come down from Pushok and Eliza. No cat in the Oliver branch is among them. the descent DBE-RE enters by a later cross, and reaches these cats Abigail Dolce Blu Crishell Dolce Blu and 3 more 5 cats stand below the collision of DBE-ALT and DBE-RE A kitten below the collision can inherit DBE-ALT, or DBE-RE, or BOTH VARIANTS IN ONE CAT, or NEITHER. Descent is not a diagnosis. Standing below two founding descents is not carrying anything, and only a test settles which of the four a given cat is. Carrying both is the compound state, and it is the one the survey records as associated with deaf and dead kittens.
Where DBE-CEL and DBE-RE meet ABOVE THE FOUNDER, WHERE THE VARIANT CAME FROM 1 animal above the DBE-CEL founders. 0 of them belong to DBE-RE. DBE-CEL and DBE-RE are not related by descent. This is the origin question, and it is closed. DBE-CEL in the genealogy: Roxi genotyped: Lazuli BELOW THE FOUNDER, WHERE THE TWO LINES MEET 130 animals below the DBE-CEL founders. 5 also have DBE-RE in the ancestry. DBE-RE arrives through the other parent in a later mating, not down through these founders. the descent DBE-RE enters by a later cross, and reaches these cats Abigail Dolce Blu Crishell Dolce Blu and 3 more 5 cats stand below the collision of DBE-CEL and DBE-RE A kitten below the collision can inherit DBE-CEL, or DBE-RE, or BOTH VARIANTS IN ONE CAT, or NEITHER. Descent is not a diagnosis. Standing below two founding descents is not carrying anything, and only a test settles which of the four a given cat is. Carrying both is the compound state, and it is the one the survey records as associated with deaf and dead kittens. Roxi is held twice in the register, as a Domestic and as a Siberian. They are one cat and the Domestic record is the fuller one.
Where DBE-ALT and the Igor line meet ABOVE THE FOUNDER, WHERE THE VARIANT CAME FROM 142 animals above the DBE-ALT founders. 0 of them belong to the Igor line. DBE-ALT and the Igor line are not related by descent. This is the origin question, and it is closed. DBE-ALT in the genealogy: Pushok, Eliza, Nikita'l Seymour, Oliver genotyped: Cotomodda Hubba-Bubba BELOW THE FOUNDER, WHERE THE TWO LINES MEET 142 animals below the DBE-ALT founders. 2 also have the Igor line in the ancestry. The Igor line arrives through the other parent in a later mating, not down through these founders. All 2 come down from Pushok and Eliza. No cat in the Oliver branch is among them. the descent The Igor line enters by a later cross, and reaches these cats Solomiya Josephine the second daughter of the two-line cross 2 cats stand below the collision of DBE-ALT and the Igor line A kitten below the collision can inherit DBE-ALT, or the Igor line, or BOTH VARIANTS IN ONE CAT, or NEITHER. Descent is not a diagnosis. Standing below two founding descents is not carrying anything, and only a test settles which of the four a given cat is. Carrying both is the compound state, and it is the one the survey records as associated with deaf and dead kittens.

A pair that can meet and has not been recorded doing so. No cat in this register stands below both, so the figure below is not a collision. It is the mating that is available inside one breed, and neither parent need show a blue eye for it to happen.

Where DBE-ALT and DBE-NAD meet ABOVE THE FOUNDER, WHERE THE VARIANT CAME FROM 142 animals above the DBE-ALT founders. 0 of them belong to DBE-NAD. DBE-ALT and DBE-NAD are not related by descent. This is the origin question, and it is closed. DBE-ALT in the genealogy: Pushok, Eliza, Nikita'l Seymour, Oliver genotyped: Cotomodda Hubba-Bubba BELOW THE FOUNDER, WHERE THE TWO LINES MEET 142 animals below the DBE-ALT founders. 0 also have DBE-NAD in the ancestry. Both are in the British Longhair and British Shorthair. The two can meet in any mating inside them. A carrier of either can show no blue eye at all, so neither parent need look like a risk. the descent DBE-NAD has not entered this descent in any recorded mating No cat in this register stands below both This pair has not met in the recorded data. It is not prevented from meeting. Nothing separates the two but the matings that have happened. A carrier of either can show no blue eye at all, so a mating that puts both variants in one kitten needs no visible warning sign, and only a test would find it afterwards.

One mating in this register brings two named variants together. On 18 July 2016, in France, Kaena, labelled DBE-ALT and recorded latent, was mated to Lazuli, DNA genotyped DBE-CEL. The litter is drawn as seven kittens on one chart and eight on another, and the published account of the same mating gives five. All three readings are held. The two deaf kittens are the one point on which every source agrees: the chart names Miyuki and Mochi and marks both with two variants, and Abitbol, Dufaure de Citres et al., Animals 2024;14:1845, records two males of this litter, one white and one almost white, both deaf. It is the only such mating the register holds.

Two variants meet in an animal only when a carrier of one is bred to a carrier of the other. A shared ancestor is a different relationship: it means two pedigrees pass through one cat, not that two alleles met in one. The shared ancestors recorded on this page were born in 2010 and 2011 and are not a meeting of alleles.

The chart labels two of the seven with both variants and those two are the two recorded deaf. Miyuki and Mochi are written DBE-CEL & DBE-ALT in red and no other kitten in the litter carries a double label. Miyuki was deaf and was not bred from. The double label is the compiler's statement of a compound heterozygote. Neither cat has been tested.

26 animals could carry two variants on this reading: the 8 of the litter and the 18 recorded below them, across 4 breeds and 2 cattery prefixes. 12 show a blue or odd eye. 26 of the 26 have never been tested for anything.

CatGenerationEyeBreed
Maidi, printed on the chart as Maidi with a diaeresisthe litter itselfunknownCeleste line
Matchathe litter itselfunknownCeleste line
Mavithe litter itselfblue-unattributedCeleste line
Mavi of Aerlin d'Aerlinthe litter itselfblue-unattributedCeleste
Merlinthe litter itselfblue-variantSiberian
Mistralthe litter itselfblue-variantCeleste line
Miyuki of Aerlin, given on the Celeste charts as Miyuki d'Aerlinthe litter itselfblue-variantCeleste line
Mochithe litter itselfblue-variantCeleste line
Aaron vom Leineuferbelow itunknownGerman Longhair
Azzurra Legende des Alpesbelow itblue-variantSiberian
Bam Bambelow itblue-variantSiberian
Ciottolina Legende des Alpesbelow itblue-variantSiberian
Conrad vom Leineuferbelow itunknownGerman Longhair
Faris vom Leineuferbelow itunknownGerman Longhair
Hani vom Leineuferbelow itunknownGerman Longhair
Jana vom Leineuferbelow itunknownGerman Longhair
Maggie vom Leineuferbelow itunknownGerman Longhair
Morgana Legende des Alpesbelow itblue-variantSiberian
Nauibelow itnot-blueCeleste line
Navajobelow itnot-blueCeleste line
Nemobelow itnot-blueCeleste line
Nessiebelow itnot-blueCeleste line
Nessybelow itodd-unattributedCeleste line
Norwinbelow itnot-blueCeleste line
Nuvolabelow itodd-variantSiberian
Tara vom Leineuferbelow itunknownGerman Longhair

What this list is and is not. It is the population in which a compound heterozygote is possible, computed from recorded parentage. It is not a list of cats that carry two variants. Each kitten of that litter took one allele from each parent at PAX3, so each carries DBE-CEL, or DBE-ALT, or one of each, and nothing but a test distinguishes the three. Below the litter the proportion falls with every outcross.

Which DBE-CEL lines could also hold DBE-ALT

117 cats of the DBE-CEL population descend from a mating that held DBE-ALT on one side, and they fall into 8 named lines. They reach that position by more than one route: one is Kaena over Lazuli of 18 July 2016, and another is a separate outcross of Roxi into the Maine Coon, which is why the Markiss cats appear here without descending from the 2016 litter. None of the 117 has been genotyped.

LineBreedsCattery prefixesCatsDNA
Blue Eyed Maine Coons (Markiss)Domestic, Experimental Longhair, Maine CoonBlue Ridge, Dolce Blu, Hanfors world, Hippopotamia, Nikita'l, Solar Eclipce47none
the Celeste line itselfCeleste, Celeste line, Maine Coon, Maine Coon variant, XLHArst, Blue Ridge, Dolce Blu, Hanfors world, d'Aerlin32none
Siberian, Altai or Barnaul unconfirmedNeva Masquerade, SiberianCoccola Siberiana, Coccole & Fusa, Great Joy, Sniewzy Urwis, Coccole & Fusa19none
German Longhair (C14)German Longhairvom Leineufer7none
Siberian (C7), from a DBE-ALT to DBE-CEL matingSiberianLegende des Alpes6none
Celeste (Chatterie d'Aerlin)British Longhair, British ShorthairChubby-Moon, d'Aerlin3none
Altai and Barnaul, both suspectedSiberianGreat Joy2none
Celeste (de Virvarnon genealogy)Celeste linenone recorded1none

A line is named here because a cat of it descends from that mating. It is not a statement that any named cat carries both variants. The lines named above are where a test would answer the question, and no cat in any of them has had one.

Every cat in which two variants could be present

Computed at every build from recorded parentage, counting both the variants the survey adopts and the ones it records a source claiming without adopting. Claimed attributions are counted here because the possibility a descendant inherits does not depend on whether the attribution met the survey's evidence threshold: Kaena's DBE-ALT is not adopted, since she was never tested, but her descendants are still in this class. 245 cats are in it.

15 of them have a determinant from each parent, which is the first generation of this population and the shortest list on the page. They are set out above under the cats with a determinant from each parent.

CatVariants that could be presentEyeBreed
Aaron Coccola SiberianaDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Aaron vom LeineuferDBE-ALT and DBE-CELunknownGerman Longhair
Abigail Dolce BluDBE-ALT and DBE-CEL and DBE-REblue-unattributedMaine Coon
Amery Dolce BluDBE-ALT and DBE-CEL and DBE-REodd-unattributedMaine Coon
Amos Coccola SiberianaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
AnaisCharm Wiking of Hanfors worldDBE-ALT and DBE-CELblue-variantMaine Coon
Angel Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedodd-variantBritish Shorthair
Anice II Dolce BluDBE-ALT and DBE-CELodd-variantMaine Coon
Aquarelle Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedlatentBritish Shorthair
Arst RobinDBE-ALT and DBE-CELblue-KITMaine Coon
Asahi Red from Sham, called Willow on the Celeste chartsDBE-CEL and Igor: PAX3, allele not publishedlatentCeleste line
Atlanta Dolce BluDBE-RE and Pillowtalk-Foreigner, unidentifiedblue-KITMaine Coon
Aurora SuperqueenDBE-ALT and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Azia Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Longhair
Azzurra Legende des AlpesDBE-ALT and DBE-CELblue-variantSiberian
Bam BamDBE-ALT and DBE-CELblue-variantSiberian
BigBoots BluraydenDBE-RE and Pillowtalk-Foreigner, unidentifiedblue-variantMaine Coon
Blue Ridge BesDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge CharizardDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge DavidDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge GalaxyDBE-ALT and DBE-CELodd-unattributedMaine Coon
Blue Ridge GavrDBE-ALT and DBE-CELblue-unattributedMaine Coon variant, XLH
Blue Ridge Gold AquamarineDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge Gregor, entered from the prefix listing as Blue Ridge GregorDBE-ALT and DBE-CELblue-KITMaine Coon
Blue Ridge GrymzaDBE-ALT and DBE-CELodd-variantMaine Coon
Blue Ridge H'Lotus PrinceDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge IolaDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge IomaDBE-ALT and DBE-CELodd-variantMaine Coon
Blue Ridge Leopold LionheartDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge Lion SpiritDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge LucienDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge Mister RedDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge MysticDBE-ALT and DBE-CELodd-variantMaine Coon
Blue Ridge NanaDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge NanaykaDBE-ALT and DBE-CELodd-variantMaine Coon
Blue Ridge Ocean DepthsDBE-ALT and DBE-CELodd-variantMaine Coon
Blue Ridge OdysseyDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge Quasar NewDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge RiksDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge RiksDBE-ALT and DBE-CELblue-KITMaine Coon
Blue Ridge RoxolanaDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge TamaraDBE-ALT and DBE-CELblue-variantExperimental Longhair
Blue Ridge U'CrushDBE-ALT and DBE-CELodd-variantMaine Coon
Blue Ridge UrbanDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge XazumDBE-ALT and DBE-CELblue-variantMaine Coon
Blue Ridge YnjirDBE-ALT and DBE-CELblue-variantExperimental Longhair
Blue Ridge YoppieDBE-ALT and DBE-CELodd-variantMaine Coon
Blue Ridge Young MaruDBE-ALT and DBE-CELblue-variantMaine Coon
Bonnie of Siberian PawsDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Briana Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantScottish Straight
Bries Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownScottish Straight
British Gems StitchDBE-ALT and Igor: PAX3, allele not publishedodd-unattributedBritish Shorthair
British Gems YumaDBE-ALT and Igor: PAX3, allele not publishedblue-unattributedBritish Shorthair
CH Quail Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Ch. Omega Great Joy, given on the Sherkan sheet as CH Omega Dreat JoyDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedblue-KITSiberian
Chery Dolce BluDBE-ALT and DBE-CELblue-variantMaine Coon
Chubby-Moon's TerrificDBE-CEL and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Ciottolina Legende des AlpesDBE-ALT and DBE-CELblue-variantSiberian
Coccole & Fusa Ilia Coccol & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedexcludedSiberian
Conrad vom LeineuferDBE-ALT and DBE-CELunknownGerman Longhair
Crishell Dolce BluDBE-ALT and DBE-CEL and DBE-REblue-KITMaine Coon
Cupcake Cinnamon SuperqueenDBE-ALT and Igor: PAX3, allele not publishedodd-variantBritish Longhair
Curly Charm Columbine from ShamDBE-ALT and Igor: PAX3, allele not publishedblue-variantSelkirk Rex
Curly Charm Lukeline Heavenly EyesDBE-ALT and Igor: PAX3, allele not publishedblue-variantSelkirk Rex
CurlyCharm SurpriseDBE-ALT and Igor: PAX3, allele not publishedunknownSelkirk Rex
Dandelion Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Diana Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Dream Dolce BluDBE-ALT and DBE-CEL and DBE-REodd-KITMaine Coon
Dream Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Elefant-Baby Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointBritish Shorthair
Elina Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
ElizaDBE-ALT and DBE-CELodd-variantDomestic
Eliza Dolce BluDBE-ALT and DBE-CEL and DBE-REodd-unattributedMaine Coon
Ermine Trace AthenaDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Ermine Trace BarkhatDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Ermine Trace JantarDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Ermine Trace LazuritDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Ermine Trace LemniscataDBE-NAD and Igor: PAX3, allele not publishedlatentBritish Longhair
Ermine Trace SapphireDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Ermine Trace UlrikaDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Ermine Trace UlrikaDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Ermine Trace XrustalDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Ermine Trace YersellaDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Excelsior Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointBritish Shorthair
Faberge Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Faris vom LeineuferDBE-ALT and DBE-CELunknownGerman Longhair
Fletcherfield's Za'MaximusDBE-RE and Pillowtalk-Foreigner, unidentifiedblue-variantMaine Coon
Flint Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Fortuna Great JoyDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Francis Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Frost Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Gessica Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointScottish Straight
Gianpaolo Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantScottish Straight
Gourami Blue Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Gourami Gold Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Gourami Pearl Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedodd-variantBritish Shorthair
Graal Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedodd-variantBritish Shorthair
Hanfors world H'SerafimDBE-ALT and DBE-CELblue-variantMaine Coon
Hanfors world H'XavierDBE-ALT and DBE-CELodd-variantMaine Coon
Hanfors world HennessyDBE-ALT and DBE-CELblue-KITMaine Coon
Hanfors World OlivkaDBE-ALT and DBE-CEL and Pillowtalk-Foreigner, unidentifiedblue-KITMaine Coon
Hani vom LeineuferDBE-ALT and DBE-CELunknownGerman Longhair
Harmony Dolce BluDBE-RE and Pillowtalk-Foreigner, unidentifiedblue-KITMaine Coon
Hatsuko Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownScottish Fold
Heavenly Charm Amigo from ShamDBE-ALT and Igor: PAX3, allele not publishedodd-variantSelkirk Rex Shorthair
Heavenly Eyes Benjamin from Sham, given on the listing as HeaventyDBE-ALT and Igor: PAX3, allele not publishedblue-unattributedBritish Shorthair
Heavenly Eyes Black Charm from ShamDBE-ALT and Igor: PAX3, allele not publishedblue-variantBritish Longhair
Hugo Perle D'Ivorie *CZDBE-ALT and a dominant blue eye, variant not named by the sourceblue-variantBritish Shorthair
Incanto Dolce BluDBE-ALT and DBE-CELblue-variantMaine Coon
Iron Coccola SiberianaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Irwin vom Wuerzburgerstern, also spelled Irven by the same sourceIgor: PAX3, allele not published and a dominant odd eye, variant not named by the sourceodd-variantBritish Shorthair
Jala Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownScottish Straight
Jana vom LeineuferDBE-ALT and DBE-CELunknownGerman Longhair
Jane Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Jessi Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Judina Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownScottish Straight
Jun Dolce BluDBE-ALT and DBE-CELblue-variantMaine Coon
King Pride AnabellDBE-ALT and Igor: PAX3, allele not publishedblue-KITBritish Longhair
King Pride ElvisDBE-ALT and Igor: PAX3, allele not publishedblue-KITBritish Shorthair
King Pride KimDBE-ALT and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
King Pride King_Pride LindaDBE-ALT and Igor: PAX3, allele not publishedlatentBritish Shorthair
King Pride MikadoDBE-ALT and Igor: PAX3, allele not publishedodd-unattributedBritish Shorthair
Kiss Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Laksheri Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointScottish Fold
Larimar Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Laser Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Lazurite Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointBritish Shorthair
Lexa Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Litter O female, Noble Teddy House, not named by the sourceIgor: PAX3, allele not published and a dominant odd eye, variant not named by the sourceblue-variantBritish Shorthair
Loena Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Lolita Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Lorenzo Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointScottish Fold
Lucilfer Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Lulluby Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Maggie vom LeineuferDBE-ALT and DBE-CELunknownGerman Longhair
Maidi, printed on the chart as Maidi with a diaeresisDBE-ALT and DBE-CELunknownCeleste line
March Nikita'l, also given as MartaDBE-ALT and DBE-CELnot-blueDomestic
Marusja DBEDBE-ALT and DBE-MARUblue-variantDomestic Shorthair
MatchaDBE-ALT and DBE-CELunknownCeleste line
Maui Dolce BluDBE-RE and Pillowtalk-Foreigner, unidentifiedblue-variantMaine Coon
MaviDBE-ALT and DBE-CELblue-unattributedCeleste line
Mavi of Aerlin d'AerlinDBE-ALT and DBE-CELblue-unattributedCeleste
Memole Dolce BluDBE-ALT and DBE-CELblue-variantMaine Coon
Meo vom OsterdeichDBE-ALT and Igor: PAX3, allele not published and a dominant blue eye, variant not named by the sourceblue-variantBritish Shorthair
MerlinDBE-ALT and DBE-CELblue-variantSiberian
Metelitsa Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Longhair
MistralDBE-ALT and DBE-CELblue-variantCeleste line
Miyuki of Aerlin, given on the Celeste charts as Miyuki d'AerlinDBE-ALT and DBE-CELblue-variantCeleste line
MochiDBE-ALT and DBE-CELblue-variantCeleste line
Morgana Legende des AlpesDBE-ALT and DBE-CELblue-variantSiberian
Morozko Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
My Mercedes Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Nao Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Narnia Dolce BluDBE-ALT and DBE-CELblue-variantMaine Coon
NauiDBE-ALT and DBE-CELnot-blueCeleste line
NavajoDBE-ALT and DBE-CELnot-blueCeleste line
Nefertiti Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedodd-variantBritish Shorthair
Neith Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
NemoDBE-ALT and DBE-CELnot-blueCeleste line
Neo Argo Chatterie de NiceDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedunknownSiberian
NessieDBE-ALT and DBE-CELnot-blueCeleste line
NessyDBE-ALT and DBE-CELodd-unattributedCeleste line
NewtDBE-CEL and Igor: PAX3, allele not publishedlatentCeleste line
NeyteriDBE-CEL and Igor: PAX3, allele not publishedunknownCeleste line
NeytiriDBE-CEL and Igor: PAX3, allele not publishedblue-variantCeleste line
Nikita'l Markiss [Cherkassy, UA]DBE-ALT and DBE-CELblue-variantExperimental Longhair
Northern Light Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Longhair
NorwinDBE-ALT and DBE-CELnot-blueCeleste line
NuageDBE-CEL and Igor: PAX3, allele not publishedblue-unattributedCeleste line
NuvolaDBE-ALT and DBE-CELodd-variantSiberian
NympheaDBE-CEL and Igor: PAX3, allele not publishedblue-unattributedCeleste line
Nyusha Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
NyxDBE-CEL and Igor: PAX3, allele not publishedblue-unattributedCeleste line
Oceania Dolce BluDBE-ALT and DBE-CELblue-variantMaine Coon
Odarka Noble Teddy House, given on the pedigree as Noble Teddy House X*OdarkaIgor: PAX3, allele not published and a dominant odd eye, variant not named by the sourceblue-KITBritish Shorthair
Omega Chatterie de NiceDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Pamela Chatterie de NiceDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Passion Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Pavel Chatterie de NiceDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Pelayo vom OsterdeichDBE-ALT and a dominant blue eye, variant not named by the sourceblue-variantBritish Shorthair
Petunia Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointBritish Shorthair
Pimprenelle D'AerlinDBE-CEL and Igor: PAX3, allele not publishedblue-variantBritish Longhair
Plamia Ermine Trace, also written Ermine Trace PlamiaDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Pollon Coccola SiberianaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Puff Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Qatar Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Longhair
Qiturah Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownScottish Straight
Quatis Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Que Guapa vom Osterdeich, written on the cattery page as Que Guapa with an acute accentDBE-ALT and a dominant blue eye, variant not named by the source and a dominant odd eye, variant not named by the sourceodd-variantBritish Shorthair
Quebec Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Quila Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Rejna Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Rhum Coda D'AerlinDBE-CEL and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Rich Great JoyDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Ricky Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Rococo Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointBritish Shorthair
Roman Great JoyDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Roxy Noble Teddy House, given by the source as J* Roxy Noble Teddy HouseIgor: PAX3, allele not published and a dominant blue eye, on an annotation that the registry sheet does not carryblue-KITBritish Shorthair
Roy Great JoyDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Santana Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Saturn Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Sherkan Great JoyDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Smile Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Sniewzy Urwis, Coccole & Fusa Ursa Sniewzy Urwis Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownSiberian
Solar Eclipce AgataDBE-ALT and DBE-CELodd-variantMaine Coon
Solar Eclipce BombeiDBE-ALT and DBE-CELblue-variantMaine Coon
Solar Eclipce FlafiDBE-ALT and DBE-CELunknownMaine Coon
Solar Eclipce IzoldaDBE-ALT and DBE-CELblue-variantMaine Coon
Solar Eclipce QueenDBE-ALT and DBE-CELblue-variantMaine Coon
Solar Eclipce ValkyriyDBE-ALT and DBE-CELunknownMaine Coon
Solomiya JosephineDBE-ALT and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
SoloMiya the second daughter of the two-line cross, not named by the sourceDBE-ALT and Igor: PAX3, allele not publishedunknownBritish Shorthair
Starfire Dolce BluDBE-ALT and DBE-CELodd-unattributedMaine Coon
Strashaya Coonka s AvitoDBE-RE and Pillowtalk-Foreigner, unidentifiedblue-variantMaine Coon
Sunny Blue Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Tanatos Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Tara vom LeineuferDBE-ALT and DBE-CELunknownGerman Longhair
Timur Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Umi Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedblue-variantSiberian
Unreal ZouWu Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Longhair
Ups Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Urania Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointBritish Shorthair
Urbi et Orbi Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedodd-variantBritish Shorthair
Ursupilami NemaususIgor: PAX3, allele not published and unnamed dominant blue eyeodd-KITBritish Shorthair
Wabisabi's JonaDBE-ALT and Igor: PAX3, allele not publishedunknownBritish Shorthair
Wispa Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
WoollyRays Iya Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantScottish Straight
X-Lily Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
X-Lotus Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
X-Water Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Xaos Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Xelia Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Xeon Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Xeres Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Xia Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedunknownNeva Masquerade
Xia Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointScottish Straight
Ximena Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Xit Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantScottish Straight
Xoxloma Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedunknownBritish Shorthair
Yellow Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedodd-variantBritish Shorthair
Yoshiki Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Zachary Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedblue-pointBritish Shorthair
Zanzibar HippopotamiaDBE-ALT and DBE-CELblue-variantMaine Coon
Zarafina Ermine Trace, also written Ermine Trace ZarafinaDBE-NAD and Igor: PAX3, allele not publishedblue-variantBritish Shorthair
Zorion Ermine TraceDBE-NAD and Igor: PAX3, allele not publishedodd-variantBritish Shorthair

The catteries in which two variants could meet

Two variants in one cattery are an ACCIDENTAL mating risk only where they sit in breeds that are bred to one another. Most catteries breed one breed, so in most cases they do. Where a cattery holds one variant in Maine Coons and another in British Shorthairs, a compound heterozygote cannot happen by accident: a mating between them would have to be deliberate. It would still be a prohibited mating. The rule this directory proposes bars carrier to carrier with no breed exception, because a breed boundary is a convention and the allele does not observe it. See the proposal.

21 red flagged, two independent variants inside one breed group. 11 black flagged, three or more inside one breed group. The rest hold their variants in separate breed groups.

DBE-RE and the Pillowtalk-Foreigner line are counted as two variants. A cat of the Pillowtalk-Foreigner line showed the dominant blue eye and was tested clear of DBE-RE. DBE-RE is nonetheless carried in Pillowtalk-descended pedigrees. Both statements hold if the line carries two alleles, since a line is not one genotype, and the register counts them separately on that reading.

The evidence on either side. For two alleles: the negative test above, on an animal the survey cannot name. For one: DBE-RE is named for a cat born in 2015 that descends from Pillowtalk cats, and Pillowtalk's Bon Aqua carries eye code 68 on a 2008 registration, seven years earlier. A variant named for a cat did not necessarily originate with that cat. One named Pillowtalk cat on a current six-variant panel would settle it, and would take six catteries off the flag list if the answer is one allele. Until then the register flags the pair, on the stated policy that an unconfirmed risk is recorded rather than dropped.

Where the branches part. Finding 3 gives the common ancestor: Love Hulen's Hurricane Mitch, MCO d 22, 18 January 1999. One son leads to Rociri's Elvis and so to DBE-RE; another son and a daughter lead to the Katrin Style cats and to Venezia the Foreigner. Mitch carries no eye code, and shared descent at that depth is common in this breed, so the split locates where to test rather than establishing an allele.

The limit of this flag. Finding 2 records that no cattery holding Rociri Elvis descent lacks Pillowtalk descent, so a flag on that pair fires on every DBE-RE cattery in the register and separates none of them. The table below therefore names the cats, the breed group and the route, so that the entry functions as a testing list. Naming the second variant would make the flag specific.

The breed grouping is a judgement of the survey and is set out so that it can be checked. The British group is wide because its breeds were derived from one another: the German Longhair and the Celeste both come from the British Shorthair, and the Azul is the breed of the Igor line. Maine Coon takes in the experimental longhair registrations; Siberian takes in the Neva Masquerade.

FlagCatteryVariants, by breedCan they meet?
BLACKBritish GemsBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair Igor: PAX3, allele not published | British Shorthair and Longhair unnamed dominant blue eyeYes, and three or more of them. British Shorthair and Longhair
BLACKCoccola SiberianaSiberian DBE, variant not named on the document | Siberian DBE-ALT | Siberian DBE-CEL | Siberian Siberian, Altai or Barnaul unconfirmedYes, and three or more of them. Siberian
BLACKCoccole & FusaNeva Masquerade DBE-ALT | Neva Masquerade DBE-CEL | Neva Masquerade Siberian, Altai or Barnaul unconfirmed | Siberian DBE-ALT | Siberian DBE-CEL | Siberian Siberian, Altai or Barnaul unconfirmedYes, and three or more of them. Neva Masquerade, Siberian
BLACKDolce BluMaine Coon DBE-ALT | Maine Coon DBE-CEL | Maine Coon DBE-RE | Maine Coon Pillowtalk-Foreigner, unidentifiedYes, and three or more of them. Maine Coon
BLACKErmine TraceBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair DBE-NAD | British Shorthair and Longhair Igor: PAX3, allele not published | Scottish Fold DBE-NAD | Scottish Fold Igor: PAX3, allele not published | Scottish Straight DBE-NAD | Scottish Straight Igor: PAX3, allele not publishedYes, and three or more of them. British Shorthair and Longhair, Scottish Fold, Scottish Straight
BLACKGreat JoySiberian DBE, variant not named on the document | Siberian DBE-ALT | Siberian DBE-CEL | Siberian Siberian, Altai or Barnaul unconfirmedYes, and three or more of them. Siberian
BLACKHanfors worldMaine Coon DBE-ALT | Maine Coon DBE-CEL | Maine Coon Pillowtalk-Foreigner, unidentifiedYes, and three or more of them. Maine Coon
BLACKNoble Teddy HouseBritish Shorthair and Longhair Igor: PAX3, allele not published | British Shorthair and Longhair a dominant blue eye, on an annotation that the registry sheet does not carry | British Shorthair and Longhair a dominant odd eye, variant not named by the sourceYes, and three or more of them. British Shorthair and Longhair
BLACKSniewzy UrwisSiberian DBE-ALT | Siberian DBE-CEL | Siberian Siberian, Altai or Barnaul unconfirmedYes, and three or more of them. Siberian
BLACKd'AerlinBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair DBE-CEL | British Shorthair and Longhair Igor: PAX3, allele not published | Celeste DBE-ALT | Celeste DBE-CEL | German Longhair DBE-CELYes, and three or more of them. British Shorthair and Longhair, Celeste
BLACKvom OsterdeichBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair Igor: PAX3, allele not published | British Shorthair and Longhair a dominant blue eye, variant not named by the source | British Shorthair and Longhair a dominant odd eye, variant not named by the sourceYes, and three or more of them. British Shorthair and Longhair
REDArstMaine Coon DBE-ALT | Maine Coon DBE-CELYes. Two independent variants sit in Maine Coon, which is one breeding group.
REDBigBootsMaine Coon DBE-RE | Maine Coon Pillowtalk-Foreigner, unidentifiedYes. Two independent variants sit in Maine Coon, which is one breeding group.
REDBlue RidgeMaine Coon DBE-ALT | Maine Coon DBE-CELYes. Two independent variants sit in Maine Coon, which is one breeding group.
REDChatterie de NiceSiberian DBE, variant not named on the document | Siberian Siberian, Altai or Barnaul unconfirmedYes. Two independent variants sit in Siberian, which is one breeding group.
REDChubby-MoonBritish Shorthair and Longhair DBE-CEL | British Shorthair and Longhair Igor: PAX3, allele not publishedYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDEmerald FlowerBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair Igor: PAX3, allele not publishedYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDFletcherfieldMaine Coon DBE-RE | Maine Coon Pillowtalk-Foreigner, unidentifiedYes. Two independent variants sit in Maine Coon, which is one breeding group.
REDHippopotamiaMaine Coon DBE-ALT | Maine Coon DBE-CELYes. Two independent variants sit in Maine Coon, which is one breeding group.
REDKing PrideBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair Igor: PAX3, allele not publishedYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDLegende des AlpesSiberian DBE-ALT | Siberian DBE-CELYes. Two independent variants sit in Siberian, which is one breeding group.
REDNemaususBritish Shorthair and Longhair Igor: PAX3, allele not published | British Shorthair and Longhair unnamed dominant blue eyeYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDNikita'lBreed not stated on the source, mostly outbred founders and first crosses Highlander and Olynx, unidentified | British Shorthair and Longhair DBE-ALT | Domestic DBE-ALT | Domestic DBE-CEL | Maine Coon DBE-ALT | Maine Coon DBE-CEL | Persian DBE-ALT | Topaz DBE-ALTYes. Two independent variants sit in Domestic, Maine Coon, which is one breeding group.
REDPerle D'IvorieBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair a dominant blue eye, variant not named by the sourceYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDSolar EclipceMaine Coon DBE-ALT | Maine Coon DBE-CELYes. Two independent variants sit in Maine Coon, which is one breeding group.
REDSoloMiyaBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair Igor: PAX3, allele not publishedYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDSuperqueenBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair Igor: PAX3, allele not publishedYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDWabisabiBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair Igor: PAX3, allele not publishedYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDfrom ShamBritish Shorthair and Longhair DBE-ALT | British Shorthair and Longhair Igor: PAX3, allele not publishedYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
REDof Siberian PawsSiberian DBE, variant not named on the document | Siberian Siberian, Altai or Barnaul unconfirmedYes. Two independent variants sit in Siberian, which is one breeding group.
REDvom LeineuferGerman Longhair DBE-ALT | German Longhair DBE-CELYes. Two independent variants sit in German Longhair, which is one breeding group.
REDvom WuerzburgersternBritish Shorthair and Longhair Igor: PAX3, allele not published | British Shorthair and Longhair a dominant odd eye, variant not named by the sourceYes. Two independent variants sit in British Shorthair and Longhair, which is one breeding group.
noneBelleAyrBritish Shorthair and Longhair DBE-ALT | Maine Coon DBE-RENo. The variants are held in separate breed groups and a mating between them would have to be deliberate.

33 cattery prefixes hold cats between them covering more than one variant, and 12 of those cover three or more. A prefix is not an animal, so each row names the cats it is computed from. Rows covering three or more are outlined. They are British Gems, Coccola Siberiana, Coccole & Fusa, Dolce Blu, Ermine Trace, Great Joy, Hanfors world, Nikita'l, Noble Teddy House, Sniewzy Urwis, d'Aerlin, vom Osterdeich.

CatteryVariants that could meet thereThe cats that put them thereCats in which both could be present
Coccola SiberianaDBE, variant not named on the document and DBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedPollon Coccola Siberiana, Amos Coccola Siberiana, Iron Coccola Siberiana, Aaron Coccola Siberiana4: Pollon Coccola Siberiana, Amos Coccola Siberiana, Iron Coccola Siberiana, Aaron Coccola Siberiana
Dolce BluDBE-ALT and DBE-CEL and DBE-RE and Pillowtalk-Foreigner, unidentifiedMaui Dolce Blu, Jun Dolce Blu, Memole Dolce Blu, Anice II Dolce Blu, Chery Dolce Blu, Narnia Dolce Blu, Oceania Dolce Blu, Incanto Dolce Blu, Abigail Dolce Blu, Atlanta Dolce Blu, Crishell Dolce Blu, Harmony Dolce Blu, Amery Dolce Blu, Dream Dolce Blu, Eliza Dolce Blu, Starfire Dolce Blu16: Maui Dolce Blu, Jun Dolce Blu, Memole Dolce Blu, Anice II Dolce Blu, Chery Dolce Blu, Narnia Dolce Blu, Oceania Dolce Blu, Incanto Dolce Blu, Abigail Dolce Blu, Atlanta Dolce Blu, Crishell Dolce Blu, Harmony Dolce Blu, Amery Dolce Blu, Dream Dolce Blu, Eliza Dolce Blu, Starfire Dolce Blu
Great JoyDBE, variant not named on the document and DBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedRich Great Joy, Roy Great Joy, Favorite Great Joy, Hunter Great Joy, Lassie Great Joy, Roxana Great Joy, Roman Great Joy, Einstein Great Joy, Sherkhan Great Joy, Yaroslava Great Joy, Yasmine Great Joy, Yalta Great Joy, Juno Great Joy, Jukka Great Joy, Upiter Great Joy, Uniq Great Joy, Unona Great Joy, Sambuka Great Joy, Skazka Great Joy, Hariton Great Joy, Hristina Great Joy, Fortuna Great Joy, Ch. Omega Great Joy, given on the Sherkan sheet as CH Omega Dreat Joy, Sherkan Great Joy6: Rich Great Joy, Roy Great Joy, Roman Great Joy, Fortuna Great Joy, Ch. Omega Great Joy, given on the Sherkan sheet as CH Omega Dreat Joy, Sherkan Great Joy
vom OsterdeichDBE-ALT and Igor: PAX3, allele not published and a dominant blue eye, variant not named by the source and a dominant odd eye, variant not named by the sourceQue Guapa vom Osterdeich, Pelayo vom Osterdeich, Que Guapa vom Osterdeich, written on the cattery page as Que Guapa with an acute accent, Meo vom Osterdeich3: Pelayo vom Osterdeich, Que Guapa vom Osterdeich, written on the cattery page as Que Guapa with an acute accent, Meo vom Osterdeich
British GemsDBE-ALT and Igor: PAX3, allele not published and unnamed dominant blue eyeBritish Gems Yuma, an unnamed British Gems cream harlequin male photographed 27 June 20221: British Gems Yuma
Coccole & FusaDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedRejna Coccole & Fusa, Neith Coccole & Fusa, Xia Coccole & Fusa, Jane Coccole & Fusa, Nao Coccole & Fusa, Lulluby Coccole & Fusa, Frost Coccole & Fusa, CH. Falco Coccole & Fusa, Umi Coccole & Fusa, Coccole & Fusa Ilia Coccol & Fusa, Jessi Coccole & Fusa, Passion Coccole & Fusa, Lexa Coccole & Fusa, Timur Coccole & Fusa, Francis Coccole & Fusa14: Rejna Coccole & Fusa, Neith Coccole & Fusa, Xia Coccole & Fusa, Jane Coccole & Fusa, Nao Coccole & Fusa, Lulluby Coccole & Fusa, Frost Coccole & Fusa, Umi Coccole & Fusa, Coccole & Fusa Ilia Coccol & Fusa, Jessi Coccole & Fusa, Passion Coccole & Fusa, Lexa Coccole & Fusa, Timur Coccole & Fusa, Francis Coccole & Fusa
Ermine TraceDBE-ALT and DBE-NAD and Igor: PAX3, allele not publishedErmine Trace Nadeya, Ermine Trace Olwen, Ermine Trace Xrustal, Ermine Trace Lemniscata, Ermine Trace Ember, Ermine Trace Athena, Azia Ermine Trace, Ermine Trace Barkhat, Ermine Trace Jantar, Ermine Trace Sapphire, Faberge Ermine Trace, Gourami Blue Ermine Trace, Quatis Ermine Trace, Quila Ermine Trace, Santana Ermine Trace, Smile Ermine Trace, Sunny Blue Ermine Trace, Tanatos Ermine Trace, Unreal ZouWu Ermine Trace, Wispa Ermine Trace, X-Lily Ermine Trace, Yoshiki Ermine Trace, Zarafina Ermine Trace, also written Ermine Trace Zarafina, Angel Ermine Trace, Ermine Trace Enya, Nefertiti Ermine Trace, Urbi et Orbi Ermine Trace, Yellow Ermine Trace, Ermine Trace Lazurit, Ermine Trace Jeana, Ermine Trace Kashemir, Ermine Trace Cherry, Manila Ermine Trace, Ermine Trace Qorall, Ermine Trace Witch, Plamia Ermine Trace, also written Ermine Trace Plamia, Ermine Trace Yersella, Saturn Ermine Trace, Ermine Trace Ulrika, Nyusha Ermine Trace, CH Quail Ermine Trace, Dream Ermine Trace, Larimar Ermine Trace, Lazurite Ermine Trace, Lolita Ermine Trace, Lucilfer Ermine Trace, Ricky Ermine Trace, Rococo Ermine Trace, Xeres Ermine Trace, Ximena Ermine Trace, Dandelion Ermine Trace, Diana Ermine Trace, Metelitsa Ermine Trace, Morozko Ermine Trace, Ups Ermine Trace, Urania Ermine Trace, Xaos Ermine Trace, Xelia Ermine Trace, Xeon Ermine Trace, Xoxloma Ermine Trace, Laser Ermine Trace, Loena Ermine Trace, My Mercedes Ermine Trace, Laksheri Ermine Trace, Lorenzo Ermine Trace, Gianpaolo Ermine Trace, Gessica Ermine Trace, Hatsuko Ermine Trace, Jala Ermine Trace, Judina Ermine Trace, Petunia Ermine Trace, Kiss Ermine Trace, Puff Ermine Trace, Qiturah Ermine Trace, Xia Ermine Trace, Xit Ermine Trace, Zachary Ermine Trace, Zorion Ermine Trace, Elefant-Baby Ermine Trace, Excelsior Ermine Trace, Briana Ermine Trace, Bries Ermine Trace, Gourami Gold Ermine Trace, Gourami Pearl Ermine Trace, Graal Ermine Trace, Flint Ermine Trace, Qatar Ermine Trace, Quebec Ermine Trace, X-Lotus Ermine Trace, X-Water Ermine Trace, Aquarelle Ermine Trace, Elina Ermine Trace, Northern Light Ermine Trace, WoollyRays Iya Ermine Trace, Ermine Trace Ulrika85: Ermine Trace Xrustal, Ermine Trace Lemniscata, Ermine Trace Athena, Azia Ermine Trace, Ermine Trace Barkhat, Ermine Trace Jantar, Ermine Trace Sapphire, Faberge Ermine Trace, Gourami Blue Ermine Trace, Quatis Ermine Trace, Quila Ermine Trace, Santana Ermine Trace, Smile Ermine Trace, Sunny Blue Ermine Trace, Tanatos Ermine Trace, Unreal ZouWu Ermine Trace, Wispa Ermine Trace, X-Lily Ermine Trace, Yoshiki Ermine Trace, Zarafina Ermine Trace, also written Ermine Trace Zarafina, Angel Ermine Trace, Nefertiti Ermine Trace, Urbi et Orbi Ermine Trace, Yellow Ermine Trace, Ermine Trace Lazurit, Plamia Ermine Trace, also written Ermine Trace Plamia, Ermine Trace Yersella, Saturn Ermine Trace, Ermine Trace Ulrika, Nyusha Ermine Trace, CH Quail Ermine Trace, Dream Ermine Trace, Larimar Ermine Trace, Lazurite Ermine Trace, Lolita Ermine Trace, Lucilfer Ermine Trace, Ricky Ermine Trace, Rococo Ermine Trace, Xeres Ermine Trace, Ximena Ermine Trace, Dandelion Ermine Trace, Diana Ermine Trace, Metelitsa Ermine Trace, Morozko Ermine Trace, Ups Ermine Trace, Urania Ermine Trace, Xaos Ermine Trace, Xelia Ermine Trace, Xeon Ermine Trace, Xoxloma Ermine Trace, Laser Ermine Trace, Loena Ermine Trace, My Mercedes Ermine Trace, Laksheri Ermine Trace, Lorenzo Ermine Trace, Gianpaolo Ermine Trace, Gessica Ermine Trace, Hatsuko Ermine Trace, Jala Ermine Trace, Judina Ermine Trace, Petunia Ermine Trace, Kiss Ermine Trace, Puff Ermine Trace, Qiturah Ermine Trace, Xia Ermine Trace, Xit Ermine Trace, Zachary Ermine Trace, Zorion Ermine Trace, Elefant-Baby Ermine Trace, Excelsior Ermine Trace, Briana Ermine Trace, Bries Ermine Trace, Gourami Gold Ermine Trace, Gourami Pearl Ermine Trace, Graal Ermine Trace, Flint Ermine Trace, Qatar Ermine Trace, Quebec Ermine Trace, X-Lotus Ermine Trace, X-Water Ermine Trace, Aquarelle Ermine Trace, Elina Ermine Trace, Northern Light Ermine Trace, WoollyRays Iya Ermine Trace, Ermine Trace Ulrika
Hanfors worldDBE-ALT and DBE-CEL and Pillowtalk-Foreigner, unidentifiedAnaisCharm Wiking of Hanfors world, Hanfors world H'Serafim, Hanfors world H'Xavier, Hanfors world Hennessy, Hanfors World Olivka5: AnaisCharm Wiking of Hanfors world, Hanfors world H'Serafim, Hanfors world H'Xavier, Hanfors world Hennessy, Hanfors World Olivka
Nikita'lDBE-ALT and DBE-CEL and Highlander and Olynx, unidentifiedNikita'l Daisy, Nikita'l Gioia, March Nikita'l, also given as Marta, Nikita'l Seymour, Nikita'l Markiss [Cherkassy, UA], Nikital Jaguar2: March Nikita'l, also given as Marta, Nikita'l Markiss [Cherkassy, UA]
Noble Teddy HouseIgor: PAX3, allele not published and a dominant blue eye, on an annotation that the registry sheet does not carry and a dominant odd eye, variant not named by the sourceJasmine Noble Teddy House, Lucia Noble Teddy House, Holly Noble Teddy House, given on the breeder chart as Mom Holly Noble Teddy House, Pallada Noble Teddy House, written on the pedigree as Noble Teddy House X*Pallada, Panda Noble Teddy House, written on the pedigree as Noble Teddy House N*Panda, Noble Teddy House Z*Unika Noble Cats, Roxy Noble Teddy House, given by the source as J* Roxy Noble Teddy House, Rosy Noble Teddy House, given on the pedigree as Noble Teddy House J*Rosy, Odarka Noble Teddy House, given on the pedigree as Noble Teddy House X*Odarka, Litter O female, Noble Teddy House, not named by the source3: Roxy Noble Teddy House, given by the source as J* Roxy Noble Teddy House, Odarka Noble Teddy House, given on the pedigree as Noble Teddy House X*Odarka, Litter O female, Noble Teddy House, not named by the source
Sniewzy UrwisDBE-ALT and DBE-CEL and Siberian, Altai or Barnaul unconfirmedSniewzy Urwis, Coccole & Fusa Ursa Sniewzy Urwis Coccole & Fusa1: Sniewzy Urwis, Coccole & Fusa Ursa Sniewzy Urwis Coccole & Fusa
d'AerlinDBE-ALT and DBE-CEL and Igor: PAX3, allele not publishedMiyuki of Aerlin, given on the Celeste charts as Miyuki d'Aerlin, Muscari of Aerlin d'Aerlin, Mavi of Aerlin d'Aerlin, Pamina d'Aerlin, Plumette d'Aerlin, Ti'Punch d'Aerlin, Urist d'Aerlin, Mirage of Aerlin d'Aerlin, Nautilus D'Aerlin, called Tilu on the de Virvarnon genealogy, Pimprenelle D'Aerlin, Rhum Coda D'Aerlin4: Miyuki of Aerlin, given on the Celeste charts as Miyuki d'Aerlin, Mavi of Aerlin d'Aerlin, Pimprenelle D'Aerlin, Rhum Coda D'Aerlin
ArstDBE-ALT and DBE-CELArst Robin1: Arst Robin
BelleAyrDBE-ALT and DBE-REBelleAyr Belle Ayr Olympas, BelleAyr Belle Ayr Onyxx, registered as GB*Belle Ayr Onyxx/FI, BelleAyr Pablo, BelleAyr Promise, BelleAyr Pauline, BelleAyr Peggy, BelleAyr Princess, BelleAyr Pluto, BelleAyr OBlue Sky, BelleAyr OBlue Sea, BelleAyr Roxanna, BelleAyr Soda Pop, BelleAyr O'Blue Pearl, BelleAyr Robin, the Maine Coon of the 23 June 2024 litter, BelleAyr Rocky, BelleAyr Russ, BelleAyr Belle Ayr O'Turner, BelleAyr Freya, BelleAyr Romulus Blue, entered here earlier as Blue, BelleAyr Rori, BelleAyr Ross Frank, entered here earlier as Ross, BelleAyr Belle Ayr Ossus, BelleAyr Robin, the British Shorthair of the 5 July 2024 litter, Belleayr Raine, BelleAyr Rio, BelleAyr Ray, BelleAyr Rivers, BelleAyr Rain, Belleayr Roxy, BelleAyr Rita, BelleAyr Rioghail Binx, BelleAyr Ronnie, BelleAyr Organanone
BigBootsDBE-RE and Pillowtalk-Foreigner, unidentifiedBigBoots Blurayden, BigBoots Belle Azure, BigBoots Big T, BigBoots Dirty Dodge1: BigBoots Blurayden
Blue RidgeDBE-ALT and DBE-CELBlue Ridge Ynjir, Blue Ridge Iola, Blue Ridge Ioma, Blue Ridge Bes, Blue Ridge Mystic, Blue Ridge Riks, Blue Ridge Nana, Blue Ridge Xazum, Blue Ridge Yoppie, Blue Ridge Young Maru, Blue Ridge Urban, Blue Ridge Lion Spirit, Blue Ridge Lucien, Blue Ridge Leopold Lionheart, Blue Ridge Grymza, Blue Ridge Gold Aquamarine, Blue Ridge Mister Red, Blue Ridge Tamara, Blue Ridge Charizard, Blue Ridge David, Blue Ridge Quasar New, Blue Ridge Nanayka, Blue Ridge Roxolana, Blue Ridge H'Lotus Prince, Blue Ridge Odyssey, Blue Ridge Ocean Depths, Blue Ridge U'Crush, Blue Ridge Gavr, Blue Ridge Galaxy, Blue Ridge Riks, Blue Ridge Gregor, entered from the prefix listing as Blue Ridge Gregor31: Blue Ridge Ynjir, Blue Ridge Iola, Blue Ridge Ioma, Blue Ridge Bes, Blue Ridge Mystic, Blue Ridge Riks, Blue Ridge Nana, Blue Ridge Xazum, Blue Ridge Yoppie, Blue Ridge Young Maru, Blue Ridge Urban, Blue Ridge Lion Spirit, Blue Ridge Lucien, Blue Ridge Leopold Lionheart, Blue Ridge Grymza, Blue Ridge Gold Aquamarine, Blue Ridge Mister Red, Blue Ridge Tamara, Blue Ridge Charizard, Blue Ridge David, Blue Ridge Quasar New, Blue Ridge Nanayka, Blue Ridge Roxolana, Blue Ridge H'Lotus Prince, Blue Ridge Odyssey, Blue Ridge Ocean Depths, Blue Ridge U'Crush, Blue Ridge Gavr, Blue Ridge Galaxy, Blue Ridge Riks, Blue Ridge Gregor, entered from the prefix listing as Blue Ridge Gregor
Chatterie de NiceDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedOmega Chatterie de Nice, Pavel Chatterie de Nice, Pamela Chatterie de Nice, Olly Chatterie de Nice, Noa Artemis Chatterie de Nice, Neo Argo Chatterie de Nice, Olivia from the Nice Cattery, held from chart material as Olivia Chatterie de Nice4: Omega Chatterie de Nice, Pavel Chatterie de Nice, Pamela Chatterie de Nice, Neo Argo Chatterie de Nice
Chubby-MoonDBE-CEL and Igor: PAX3, allele not publishedChubby-Moon's Terrific1: Chubby-Moon's Terrific
Emerald FlowerDBE-ALT and Igor: PAX3, allele not publishedWanessa Emerald Flower, Paula Emerald Flower, Ch. Libera Emerald Flower*RUnone
FletcherfieldDBE-RE and Pillowtalk-Foreigner, unidentifiedFletcherfield's Jack the Ripper, Fletcherfield's Chardonnay, Fletcherfield's Za'Xandergroots, Fletcherfield's Za'Maximus, Fletcherfield's Za'Merlin, Fletcherfield's Blauer Klaus1: Fletcherfield's Za'Maximus
HippopotamiaDBE-ALT and DBE-CELZanzibar Hippopotamia1: Zanzibar Hippopotamia
King PrideDBE-ALT and Igor: PAX3, allele not publishedKing Pride Kim, King Pride Q-Bert, King Pride Lady Tiffany, King Pride Umka, King Pride Yasmin, Tender Paws Michelangelo of King Pride, King Pride Dior, Vanilla Cleo of King Pride, King Pride Calista, King Pride Red Diamond, King Pride Red Fire, King Pride Konsul King's Pride, Efremovit Domino of King Pride, King Pride Anabell, King Pride Elvis, King Pride King_Pride Linda4: King Pride Kim, King Pride Anabell, King Pride Elvis, King Pride King_Pride Linda
Legende des AlpesDBE-ALT and DBE-CELAzzurra Legende des Alpes, Morgana Legende des Alpes, Ciottolina Legende des Alpes3: Azzurra Legende des Alpes, Morgana Legende des Alpes, Ciottolina Legende des Alpes
NemaususIgor: PAX3, allele not published and unnamed dominant blue eyeUrsupilami Nemausus1: Ursupilami Nemausus
Perle D'IvorieDBE-ALT and a dominant blue eye, variant not named by the sourceHugo Perle D'Ivorie, Felicitas Perle D'Ivorie, Hugo Perle D'Ivorie *CZ1: Hugo Perle D'Ivorie *CZ
Solar EclipceDBE-ALT and DBE-CELSolar Eclipce Agata, Solar Eclipce Bombei, Solar Eclipce Izolda, Solar Eclipce Flafi, Solar Eclipce Queen, Solar Eclipce Valkyriy6: Solar Eclipce Agata, Solar Eclipce Bombei, Solar Eclipce Izolda, Solar Eclipce Flafi, Solar Eclipce Queen, Solar Eclipce Valkyriy
SoloMiyaDBE-ALT and Igor: PAX3, allele not publishedChristian Solomiya, Solomiya Josephine, SoloMiya the second daughter of the two-line cross, not named by the source2: Solomiya Josephine, SoloMiya the second daughter of the two-line cross, not named by the source
SuperqueenDBE-ALT and Igor: PAX3, allele not publishedKooper Superqueen, Cupcake Cinnamon Superqueen, Bellinda Superqueen, Celine Superqueen, Ed Hardy Superqueen, Elizabeth SuperQueen, Fiona SuperQueen, Holly Molly Superqueen, Nika SuperQueen, Qantus Superqueen, Tansy SuperQueen, Wings SuperQueen, Winjs SuperQueen, Y-Kira SuperQueen, Ziata SuperQueen, Aura SuperQueen, Superqueen Holms Super Queen1: Cupcake Cinnamon Superqueen
WabisabiDBE-ALT and Igor: PAX3, allele not publishedWabisabi's My Mona-Lee, Wabisabi's Jona1: Wabisabi's Jona
from ShamDBE-ALT and Igor: PAX3, allele not publishedHeavenly Eyes Ultra Black from Sham, Heavenly Eyes Benjamin from Sham, given on the listing as Heaventy, Pierce Brosnan from Sham, Ricco Fawny from Sham, Rosalie Felicitas from Sham1: Heavenly Eyes Benjamin from Sham, given on the listing as Heaventy
of Siberian PawsDBE, variant not named on the document and Siberian, Altai or Barnaul unconfirmedBonnie of Siberian Paws1: Bonnie of Siberian Paws
vom LeineuferDBE-ALT and DBE-CELJana vom Leineufer, Aaron vom Leineufer, Faris vom Leineufer, Conrad vom Leineufer, Hani vom Leineufer, Maggie vom Leineufer, Tara vom Leineufer7: Jana vom Leineufer, Aaron vom Leineufer, Faris vom Leineufer, Conrad vom Leineufer, Hani vom Leineufer, Maggie vom Leineufer, Tara vom Leineufer
vom WuerzburgersternIgor: PAX3, allele not published and a dominant odd eye, variant not named by the sourceLaurina vom Wuerzburgerstern, Lolita vom Wuerzburgerstern, Jocker vom Wuerzburgerstern, posted as Jocker vom Wuerzbgerstern, Unnamed male, ny 11 08 61, born 5 May 2024 vom Wuerzburgerstern, Unnamed male, ny 11 61, born 15 July 2024 vom Wuerzburgerstern, Girl 1, ny 12 61, first blue-eyed litter vom Wuerzburgerstern, Girl 2, ny 11 61, odd-eyed, first blue-eyed litter vom Wuerzburgerstern, Girl 3, by 12 61, first blue-eyed litter vom Wuerzburgerstern, Boy 1, ny 12 61, first blue-eyed litter vom Wuerzburgerstern, Barby vom Wuerzburgerstern, Karina vom Wuerzburgerstern, Irwin vom Wuerzburgerstern, also spelled Irven by the same source, Jessi vom Wuerzburgerstern1: Irwin vom Wuerzburgerstern, also spelled Irven by the same source

The risk is a property of the mating, not of the individual. A latent carrier of one variant bred to a carrier of another produces the compound heterozygote, and a latent carrier is not identifiable on inspection: Kaena is recorded with light green eyes and a white locket. That is the case these tables are computed to locate.

The unidentified variants

How they are named here. A variant has no official name until it is confirmed by genotype. Each row below is therefore written under the full name of the line it is possibly associated with, and nothing else: Pillowtalk-Foreigner, Igor, Karagaisky, Nadeya. The name is a label for a line, not a designation for an allele.

Unidentified variantRecordsBlue or oddPrefixesBelowAboveCollateralDNA
Igor: PAX3, allele not published400716083808817none
Blue Eyed Maine Coons (Markiss), DBE-CEL by test on the generation below98541915110695none
Siberian, Altai or Barnaul unconfirmed, unidentified1305135366624none
Topaz foundation (C6 and C11), unidentified5720715134307none
Karagaisky, unidentified18142770none
Highlander, Olynx (C11), unidentified1052360none
King's Way, unidentified752000none
Pillowtalk-Foreigner, unidentified61318262128none
Royal Edelweiss, unidentified211196286none
Gosha, unidentified100000none

What this table is not. It is not a claim that these are distinct alleles. Two rows could be the same variant, or one could be a variant already named. The grouping follows the register's own line labels and nothing else, merged only where those labels are one programme written several ways. The DNA column is empty down its whole length.

The blue eyes no variant accounts for

538 cats in this register show a blue or an odd eye. 130 of them carry a named variant. The other 408 do not, and they divide in two.

313 stand on a named line. Of those, 214 stand on a line for which no variant has been identified at all: there are 32 such lines, holding 1069 records between them, and not one animal in any of them has been genotyped. Each is a documented breeding programme with founders, dates and cattery prefixes, and no allele named.

95 more stand on no line at all, having neither traced descent nor a test. This is the largest single class in the register, and 105 of the unaccounted-for are British Shorthair.

LineRecordsBlue or oddCattery prefixes
Igor4007160
Siberian, Altai or Barnaul unconfirmed953632
Malvina / Barnaul32153
Mink and pointed Ragdoll (C9), not a DBE line36153
Karagaisky18142
Topaz foundation (C6 and C11)18112
Altai / Nikita'l Seymour2892
DBE-CEL Sphynx (C5)57727
King's Way752
Highlander, Olynx (C11)1052
Malvina / Barnaul or Aktau / Caspian, not resolved441
Kotomodda, by descent from Hugo Kotomodda442
Pillowtalk-Foreigner61318
Celeste (breeder sites)533
Superqueen432
Pedigree ancestry, not a DBE line2492130
Siberian (Chatterie de Nice descent)222
Aktau / Caspian110
Royal Edelweiss211
Seymour, by the from Sham cattery's own chart111
Seymour110
possibly SoloMiya, on a coincidence of codes only111

One of these lines has been tested and came back negative. A cat of the Pillowtalk-Foreigner line showed the dominant blue eye and was clear of DBE-ALT, DBE-RE and DBE-CEL. THAT PANEL WAS THE THREE VARIANTS DESCRIBED AT THE TIME. DBE-AGO, DBE-cDc and DBE-xHW were named afterwards, so the result reads as not one of the three then known and not as a ninth variant. DBE-AGO in particular is a later-described Maine Coon variant the panel could not have covered. Retesting one cat of that line against a current panel would either confirm a ninth variant or resolve the line to DBE-AGO.

Coverage. The eight identified variants account for 130 of the 538 cats recorded here as showing a blue or an odd eye, or 24 per cent.

Tracing them

A cattery name is not a unit of inheritance. This page ignores the prefix field entirely and traces each variant through recorded parent links and nothing else.

How a dominant allele is traced, and what each column counts

A dominant allele comes down from an ancestor. The animals that could carry it are therefore the descendants of any ancestor of a known carrier. Three tiers are computed and kept apart, because they are not equally strong. Where a variant has a laboratory result anywhere, the trace is anchored on the genotyped animals only.

Attributed
Records to which some source gives this variant, at any evidence grade. A cat is counted here whether it was DNA genotyped, fixed by a first-degree genotype, traced to a named founder, or simply asserted on a breeder's chart. It is not a count of tested cats.
DNA
Of those, how many hold a laboratory result for this animal. This is the only column that rests on a test.
Below
Recorded descendants of the anchor animals. A cat here that shows the eye must have had it from the anchor.
Above
Recorded ancestors of the anchor animals. One of them transmitted it; which one is a separate question.
Collateral
The other descendants of those ancestors. These could carry it and almost none has been tested. This is the class the testing list is drawn from.
Shares ancestors with the Igor chart
Whether any animal in the three tiers stands on the breeder-compiled chart headed line descended from Altai gene Igor. It is a statement of shared ancestry and not of direction, and nothing here says whether the shared ancestors carried a variant at all.
VariantAttributedDNABelowAboveCollateralShares ancestors with the Igor chart
DBE-ALT481622448484yes, 26 records
DBE-NAD42191154267yes, 18 records
DBE-RE301214173187no
DBE-CEL24171259no
DBE-MARU51400no
DBE-xHW40000no
DBE-cDc20000no
DBE-AGO10000no

The one variant that shares ancestors with the Igor chart

DBE-ALT, through 210 shared ancestors: Abdullah Vom Haus Aden, Ambrosius V Domus Felium, Amor van Wier en Gouwe, Antony, Benny von Buergersruh, Big Platschis Bernie Plumperquatsch, CH Artos von Quintana, CH Carajan zur Sternebucht, CH Dellary's Exoti Merlin, CH Du Blue D'Or Argentin Nuance's Golden Quantin, CH Golden-Hironimus vom Goldenen Winkel, CH Golden-Ivanhoe vom Goldenen Winkel, CH High Hesket's Silver O'Connor, CH Idefix von Konradsburg, CH It's Me Ior Van Elsdieck, CH Marvellous Silver Ramzes II, CH Silver-Filou von Wahrberg, CH Sir Eddy von Elami's, Captain Ermine Trace, Chatfou Minton, Du Blue D'Or Argentin Nuance's Golden Gotto, EC Annaida's Enrico Caruso, EC Asterix of Sarsen Stones, EC Darius-Doethe V Edelfeld, EC Gold-Amaretto D'Oaxaca, EC Golden-Erasmus von Wahrberg, EC Jadwiga's Silver Don Quichotte, EC Maurice vom Wahrberg, EC Quandili Quicks De Tuatha De Danann, EC Silver Valentyn van Elsdieck, Elvis of Sarsen Stones, Eric von Buergersruh, given on the sheet as Eric von Bürgersruh, Esteban Von Aemestelhove, Extasy's Cashman of Golden Flame, Florah's Theodoor, Flying Magic Blue Santiago of Golden Flame, GC Pennydown Top Gunn, GEC Quentin of Mystery's Dream, GIC Andy vom Wahrberg, GIC Fernando van Ybloske-Eksolby, GIC Marvellous Golden Radetzky, GIC Silver-Gary V Quintana, GIC Sir-Xenon vom Goldenen Winkel, GIC Sulver Romeo fan Liwardia, Golden Aljoscha von der Kraepelinburg, Golden-Avalon Ahawa, Golden-Gigolo, Golden-Jappeloup D'Oaxaca, Goldstern De Broeckloni, Grandeur's G. Boy I. Holland, IC Bee-Jay Golden Macho Boy of Golden Flame, IC Eddie vom Kreuzanger, IC El Bimbo of Sissinghurst, IC Freedom zur Raeuberhoehle, IC Golden-Toby von Wahrberg, IC Goldfever's Golden Mikka, IC Marvellous Masterplayer, IC Sir Quincy von Elami's, IC Sulver Cupido fan Liwardia, Insider from Sham, Jadwiga's Karate Kid, Joris, Judifax Joldran, Judifax Justus, Marvellous Golden Deister, Marvellous Golden Glorieux, Marvellous Silver Prince, Miramar Master Elrond, Murphy von Shoka-Thai, Muslin Cats Patrick Peach, Nando v Waerinveste, Nanook's Silver Ascot, Oliver-Shin Boray v't Grashus, Patchouli De Tuatha-De-Danann, Peerless Ice Breaker, Peerless Silver Monarch, Polar Jenko, Potentilla's Stoljar Stanislav, Potentilla's Zdzislav Zbigniew, Prometheus V Aemstelveste, Quassi Felis Pretiosus, Roman Aus Der Harzerland, Rosewood vum Rosegaard, Ruedisueli's Xoebiedoe Sjefke, Sekmet's Silver Sinian, Silver Jojo from Sunnyplace, Silver Kotek of Jadwiga's Home, Silver-Asterix vom Bleek, Silver-Brend Fan Llwardia, Silver-Esteban D'Oaxaca, Sonnyboy Zur Porta Westfalica, Sweet Heartbreaker's Love Me Tender, Tjarden von Felden, Valiant vh Ridderhof, WCH Bluegold Nougat D'Oaxaca, WCH Greendale Sargeant Pepper, Yeti Aus Dem Harzenland, Zinedin of Mystery's Dream, Zur Sternenbucht Golden Macavity, Amanda, given on the sheet as Amanda (D of Roman V Harzerland), Anchjin from Dust of Silver, Ariadne v Tyolet, Arminia Felis Pretiousus, Assia V. Bocca Concolor, Babettchen Vom Schloss Bordelum, Beauty Anak Dari Bella, Beauty Van Beverwijk, Beauty's Lois van Beverwijk, Big Platschis Ivi Turbosocke, CH Belia v Haus Fasler, CH Cora Dominika Peppercats, CH Di Parseghian's Vicky, CH Du Blue D'or Argentin Nuance's Golden, CH Gloria of Snow Queen, CH Golden Uglaya vom Juwelental, CH Jadwiga's Golden Undina, CH Kandy von Romero, CH Lady Di von Wahrberg, CH Lady Uliah von Elami's, CH Nicol Snow Queen, CH Noortje van de Pronkse Schonen, CH Ondra von Ursus Minor, CH Pennydown Katie Starlight, CH Philippa vom Juwelental, CH Silver Emmely from Sunnyplace, CH Silver-Whitney v Hileira, Cheeky, Clarinda Zolotoi Zamok, Cosima of Sarsen Stones, Danaclara of Sissinghurst, Du Blue D'Or Argentin Nuance's Golden Juana, EC Alice of Golden Beauty, EC Annaida's Silver Barberella, EC Ennypenny Fan Anjulies Balloo, EC Gipsy of New Celtic Henge, EC Golden-Willow von Wahrberg, EC Silver Venus of Jadwiga's Home, EC Wilma-point von Wahrberg, Farah Boroy v't Grashus, Florence Rejinald, GIC Annaida's Golden Hathly, GIC High Hesket's Silver Ivory, GIC Jadwiga's Golden Rosamunde, GIC Patty von Felden, GIC Petunia Snow Queen, GIC Silver Coralie von der Anger, GIC Sulver Querina Fan Liwardia, Gaya of Pointdreams, Gina of Sarsen Stones, Golden Debby vom Rosentor, Golden Flame vom Juwelental, IC Lady Bernice vom Goldenen Winkel, IC Marvellous Miss Goldfever, IC Queen of Beauty Peppercats, IC Silver Amy-Lee from Sunnyplace, IC Silver Clara vom Bleek, IC Silver-Nadine von Wahrberg, IC Yara von Felden, J'taime Aleeshane of Golden Flame, Jadwiga's Silver Classic Touch, Jadwiga's Silver Dollar Girl, Jadwiga's Silver Primadonna, Janosch's Chinoa, Josefine of Sarsen Stones, Joy von Romero, Judifax Jacha, Juniors Rowena, Kaori de Maneki Neko, L'Avvenires Deanna Troi, Lady Pinkybell von Elami's, Lirazel De Tuatha-De-Danann, Lizarel De Tuatha De Danann, Marcutt White Unique, Marvellous Golden Desiree, Marvellous Golden Dream, Marvellous Golden Orchidee, Marvellous Golden Samba, Marvellous S. Priceless, Miuchi de Maneki Neko, Muslin Cats Bonita Bice, Nahimana's Leilani, Nina Felis Pretiosus, Oshilla De Tuatha-De-Danann, Pearl of Empire's High Hope, Pearl of Empire's Jaikie, Peerless Marvellous Ria, Peerless Silver Sparkle, Pemberly's Silver Kassandra, Potentilla's Dewotsjka Djalixa, Potentilla's Fenja-Fjodora, Potentilla's Xorosjii, Potentilla's Zosislawa Zuzowa, Quendula von Felden, Semiramis von Felden, Shirley Anak Dari Bella, Silver Dunja of Jadwiga's Home, Silver Irina of Jadwiga's Home, Silver Nina von Tonkay, Silver Oberta Fan Liwardia, Silver-Carrie von Quintana, Silver-Donna von Quintana, Silver-Fay v Quintana, Silver-Sheba Vd Oldehove, Silvermoon's Samira, Silvester vd Oldehove, Tieni von Wahrberg, Tilly von Westhausen, Tyolet's Kim, WCH Carina vom Kreuzanger, Za-Za of New Celtic Henge. Two of them are the halves of a single mating. Every step is a recorded parent and no cattery name is used at any point. Finding 23 sets it out animal by animal.

Two branches off one mating, four generations down each side. That is a collateral relationship and it carries no direction. The shared pair is also too young to be the origin of DBE-ALT: Zinedin of Mystery's Dream was born 7 August 2011, and the variant is named for the Altai and traces to Fyodor in Kazakhstan in 1995 or 1996. Neither animal of the shared pair has been tested, so whether either carried a variant is not recorded.

The animal it ends on is Barbariska Elisa Vetsky, four generations below the shared pair. She is green-eyed, on the compiler's own knowledge of the cattery, so the chain does not end in a cat showing the trait; her blue-eyed kittens are by a sire who is coded 61 and descends from Igor. A PAX3 panel on one of those kittens, not on her, is what would say whether the Igor blue eye is DBE-ALT or a ninth variant.

What the table shows about the rest

DBE-CEL traces upward through one animal. Its one genotyped cat, Lazuli, is drawn on every chart out of an unnamed female by Roxi. The line holds 2 ancestors and 59 collateral relatives, all of them reached through that one parent link, and Roxi is computed lineage genotyped DBE-CEL: his son carries it, a dominant allele must come from a parent, and the dam is drawn as a question mark.

A trace reports the record, not the cats. Every figure on this page is a floor computed at the depth the register currently holds, and a single missing parent link is enough to make a line read as unreachable.

DBE-RE has the deepest ancestry of any variant here and reaches nothing. 173 ancestors above its genotyped animals and 187 collateral, and not one touches the Igor chart. Depth of pedigree is not what decides whether two lines meet.

No cat in the survey holds a verified genotype test showing two identified variants. A verified genotype test means a laboratory result for a named variant, on a named animal, which the survey has seen. 31 animals here hold a genotype result, and each result names one variant and stands on a different cat. The results were not commissioned or verified by the survey: they were supplied by owners or taken from work already published. Any statement that two of them meet in one cat rests on pedigree, not on a verified genotype test.

The pedigrees say the same. Five of the eight identified PAX3 variants have a carrier in this register: DBE-AGO, DBE-ALT, DBE-CEL, DBE-MARU, DBE-NAD, DBE-RE, DBE-cDc, DBE-xHW. Traced from their genotyped animals and tested as 28 pairs, no two of their pedigree webs meet at a single animal. Every meeting the survey has found between blue-eyed populations is a meeting with the Igor variant, which is at PAX3 with its allele not published, named after the cat Igor Azure Dream, and not a meeting between two identified variants.

Traced from every attributed carrier instead of from the genotyped animals only, two pairs do meet. DBE-ALT and DBE-CEL hold ancestors in common, and so do DBE-ALT and DBE-RE. Those traces run through cats that were given a variant by a chart or by descent rather than by a laboratory, so the meeting is a meeting of attributions and not of test results. The grid in the directory sets out every pairing on that wider basis.